Conspectus cobitidum - Raffles Museum of Biodiversity Research

ures copied from earlier works and not representative of the
concerned populations.
It seems that a group of populations have a short and
tapering caudal peduncle with high dorsal and ventral crests
originating immediately behind dorsal and anal fins, a deep
body and a somewhat pointed caudal fin. This agrees with
the holotype of P. dabryanus (figured by Fang, 1935b: fig. 11)
and possibly M. oligolepos. The original description of
M. mizolepis also mentions the high crests on the caudal peduncle.
Chen (1981) placed M. mizolepis in the synonymy
of P. dabryanus.
A second group has a more slender body, shorter and
less deep crests and a more rounded caudal fin, which has
often been called M. mizolepis. This may correspond to several
of the subspecies of M. mizolepis described by Nichols
and others and which Chen (1981) placed in the synonymy
of M. anguillicaudatus. This is tentatively followed here but
clearly it should be re-evaluated after examination of more
material and the the type material of all.
Nomenclature notes. The earliest description of M. oligolepos
as a new species that I could examine is by Chen,
Shen & Li (1994). Wang et al. (2001: 179) referred to a paper
by Li (1993a), which could not be found. There are also
mentions of the description of a Misgurnus olygoskolopos
in a paper by Li (1993b), which also cannot be found (Chen
Xiao-Yong, pers. comm.).
4.19 Protocobitis Yang & Chen, 1993
Protocobitis Yang & Chen, 1993: 125 (type species: Protocobitis
typhlops Yang & Chen, 1993: 125, by monotypy;
also in Yang, Chen & Lan, 1994: 92). Gender feminine.
Taxonomic notes. It remains to be demonstrated that the
two cave species P. polylepis and P. typhlops are really congerenic,
that is, that they share an immediate common ancestor.
This implies either that the ancestor was an epigean
species and is now missing or still uncollected, or that this
ancestor was a cave species that colonised two disjunct cave
areas. Each species likely has a surface ancestor.
Nomenclatural notes. Protocobitis and P. typhlops, both
by Yang & Chen (1993), predate the same names in Yang et
al. (1994) (Kottelat & Bréhier, 1999: 356).
4.19.1 'Protocobitis' polylepis Zhu, Lü, Yang & Zhang, 2008
Protocobitis polylepis Zhu, Lü, Yang & Zhang, 2008: 453,
fig. 1 (type locality: China: Guangxi: Wuming County:
underground source, 5 km from Wuming, 23°10'N
108°17'E; holotype: KIZ 20060001; compound noun,
indeclinable)
4.19.2 Protocobitis typhlops Yang & Chen, 1993
Protocobitis typhlops Yang & Chen, 1993: 125 (type locality:
China: Guangxi: Du'an County: Xia'ao town, cave,
24°15'N 107°05'E; lectotype: KIZ 918001, designated
by Kottelat & Bréhier, 1999: 356; also in Yang, Chen &
Lan, 1994: 93; compound noun, indeclinable)
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4.20 Sabanejewia Vladykov, 1829
Sabanejewia Vladykov, 1929: 86 (type species: Cobitis balcanica
Karaman, 1922: 307, by original designation).
Gender feminine.
4.20.1 Sabanejewia aurata (Filippi, 1863)
Cobitis aurata Filippi, 1863: 391 (type locality: Iran: stream
near Sartschem [apparently Sarcham-e Sofla, 39°07'N
47°54'E, B. W. Coad, pers. comm.]; lectotype: MZUT
674, designated by Tortonese, 1961: 188; adjective, -us,
-a, -um)
Cobitis hohenackeri Kessler, 1877: 177 (type locality: Azerbaidjan:
Kura River, Caspian Sea basin [Berg, 1949: 894];
syntypes: ZISP 2854 [5], Eschmeyer & Fricke, 2010;
author indicated as Brandt, but Kessler is actual author;
noun in genitive, indeclinable)
? Cobitis aralensis Kessler, 1877: 184 (type locality: Kazakhstan:
mouth of Syr-Darya River / Uzbekistan: mouth
of Amu-Darya River [Berg, 1949: 895]; syntypes [5]:
ZISP 10932 [1], Eschmeyer & Fricke, 2010; adjective,
-is, -is, -e)
4.20.2 Sabanejewia balcanica (Karaman, 1922)
Cobitis balcanica Karaman, 1922: 307 (type locality: Serbia:
Save River near Zagreb / Bosnia-Herzegovina: Miljacka
Stream, tributary of Bosna / FYROM: Vardar River
(that is, its tributaries near Skopje (Üskub) and Veles)
[type locality restricted by Bănărescu et al., 1972: 1 but
irrelevant as this can only be done by lectotype designation];
syntypes: MMNHS lost, ? MNHN 28.222 [1], Bertin
& Estève, 1948: 93; adjective, -us, -a, -um)
Cobitis montana Vladykov, 1925: 320, pl. 8 (type locality:
Ukraine: streams Apica, Tereovka, Teresulka, Luzanka
and Terebla, tributaries of Theiss River [Tisza] / Theiss
River between villages Akna-Slatina and Butino; syntypes:
[LU] 219; junior primary homonym of Cobitis
montanus Jerdon, 1849: 332; adjective, -us, -a, -um)
Cobitis aurata balcanica natio radnensis Jaszfalusi, 1951:
116, pl. 1 fig. 1 (infrasubspecific, name not available;
locality: Romania: Zebrak River, a tributary of Szamos
River [Somesul ?] / Maros River [Mures] near Gödemesterhaza
and creek Göde)
Cobitis aurata balcanica morpha elongata Oliva, Balon &
Franck, 1952: 295 (infrasubspecific, name not available;
locality: Czechia: Moravia: Lipník on Bečva River)
Cobitis aurata balcanica morpha elata Oliva, Balon &
Franck, 1952: 295 (infrasubspecific, name not available;
locality: Czechia: Moravia: Lipník on Bečva River)
Cobitis aurata radnensis Bănărescu, Müller & Nalbant,
1960: 124 (type locality: Romania: upper Mures and Tirnava
Rivers; syntypes: LU; adjective, -is, -is, -e)
Cobitis aurata bosniaca Karaman, 1963: 629, fig. 1 (type
locality: Bosnia-Herzegovina: creeks Saturlija and Siroka
near Banja Luka, Vrba River drainage; syntypes [144]:
MMNHS, lost; adjective, -us, -a, -um)
? Sabanejewia aurata doiranica Economidis, 1995: 211
(nomen nudum)
? Sabanejewia aurata thrakica Economidis, 1995: 211
(nomen nudum)
? Sabanejewia aurata doiranica Economidis & Nalbant,