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Conspectus cobitidum - Raffles Museum of Biodiversity Research

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leptes on p. 311, first reviser [Nalbant, 1963: 364] gave<br />

precedence to Somileptus, see Kottelat, 1998c: 118).<br />

Gender masculine.<br />

4.4.1 Canthophrys gongota (Hamilton, 1822)<br />

Cobitis gongota Hamilton, 1822: 351, 394 (type locality:<br />

India: "rivers <strong>of</strong> Northern Bengal towards the mountains"<br />

[Bangladesh: Patgong, Rangpur District, 25°45'N<br />

89°15'E; Hora, 1935a: 49]; types: lost; Hamilton's unpublished<br />

figure reproduced in M'Clelland, 1839: pl. 51<br />

fig. 1; noun in apposition, indeclinable)<br />

Cobitis cucura Hamilton, 1822: 352, 394 (type locality: India:<br />

Kosi River [at Nathpur near the Nepal frontier; Hora,<br />

1935a: 49]; types: lost; Hamilton's unpublished figure<br />

reproduced in M'Clelland, 1839: pl. 51 fig. 2; simultaneous<br />

subjective synonym <strong>of</strong> Cobitis gongota Hamilton,<br />

1822: 351, first reviser [apparently Günther, 1868: 363]<br />

gave precedence to C. gongota; noun in apposition, indeclinable)<br />

Cobitis oculata M'Clelland, 1839: 303, 433, pl. 51 fig. 1<br />

(type locality: India: Upper Assam / northern parts <strong>of</strong><br />

Bengal near the foot <strong>of</strong> the mountains; syntypes: SMF<br />

412 [4], 8979 [1], Eschmeyer & Fricke, 2010 [plus basis<br />

<strong>of</strong> Cobitis gongota Hamilton, 1822: 351]; adjective, -us,<br />

-a, -um)<br />

Canthophrys albescens Swainson, 1839: 310 (available by<br />

indication to Hamilton, 1822: "Cob. No. 3" [which is<br />

Cobitis cucura]; type locality: India: Kosi River [at Nathpur<br />

near the Nepal frontier; Hora, 1935a: 49]; types: NT;<br />

Hamilton's unpublished figure reproduced in M'Clelland,<br />

1839: pl. 51 fig. 2; participle, indeclinable)<br />

Somileptes bispinosa Swainson, 1839: 311 (available by indication<br />

to Hamilton, 1822: 351 [which is Cobitis gongota];<br />

type locality: India: "rivers <strong>of</strong> Northern Bengal towards<br />

the mountains" [Bangladesh: Patgong, Rangpur<br />

District, 25°45'N 89°15'E; Hora, 1935a: 49]; types: lost;<br />

Hamilton's unpublished figure reproduced in M'Clelland,<br />

1839: pl. 51 fig. 1; adjective, -us, -a, -um)<br />

? Cobitis amnicola Valenciennes, in Cuvier & Valenciennes,<br />

1846: 68 (type locality: India: Bengal; syntypes: MNHN<br />

B.2171 [3], Spillmann, 1962: 587; noun in apposition,<br />

indeclinable)<br />

4.5 Cobitis Linnaeus, 1758<br />

Cobitis Linnaeus, 1758: 303 (type species: Cobitis taenia<br />

Linnaeus, 1758: 303, designated by ICZN, 1988 [Opinion<br />

1500]). Gender feminine.<br />

Acantophthalmus van Hasselt, 1823:132 (type species: Cobitis<br />

taenia Linnaeus, 1758: 303, by monotypy [see Kottelat,<br />

1987b: 371]; on Official Index <strong>of</strong> Rejected Names,<br />

ICZN, 1988: 178 [Opinion 1500]). Gender masculine.<br />

Acanthophthalmus van Hasselt, 1824: 377 (incorrect subsequent<br />

spelling <strong>of</strong> Acantophthalmus van Hasselt, 1823:<br />

132; on Official Index <strong>of</strong> Rejected Names, ICZN, 1992b:<br />

248 [Opinion 1695])<br />

? Acanthopsis Agassiz, 1832: 134 (type species: Acanthopsis<br />

angustus Agassiz, 1835: vol. 5, pl. 50 fig. 2, by subsequent<br />

monotypy in Agassiz, 1835: pl. 50; not a junior<br />

homonym <strong>of</strong> Acanthopsis van Hasselt, 1824: 377, which<br />

THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />

23<br />

is an incorrect subsequent spelling <strong>of</strong> Acantopsis van<br />

Hasselt, 1823: 133, thus does not enter into homonymy<br />

[Code art. 33.3]; based on a fossil species, possibly not a<br />

cobitid (Kottelat, 1987b: 372)). Gender feminine.<br />

Cobitinula Hankó, 1924: 152 (type species: Cobitinula anatoliae<br />

Hankó, 1924: 152, by monotypy). Gender feminine.<br />

Acanestrinia Băcescu, 1962b: 435 (subgenus <strong>of</strong> Cobitis Linnaeus,<br />

1758: 303; type species: Cobitis elongata Heckel &<br />

Kner, 1858: 305, by original designation). Gender feminine.<br />

Bicanestrinia Băcescu, 1962b: 436 (subgenus <strong>of</strong> Cobitis<br />

Linnaeus, 1758: 303; type species: Cobitis simplicispina<br />

Hankó, 1924: 153, by original designation). Gender<br />

feminine.<br />

Iberocobitis Băcescu, 1962b: 438 (subgenus <strong>of</strong> Cobitis Linnaeus,<br />

1758: 303; type species: Acanthopsis taenia paludica<br />

de Buen, 1930: 34, by original designation). Gender<br />

feminine.<br />

Beyshehiria Erk'akan, Atalay-Ekmekçi & Nalbant, 1999: 20<br />

(subgenus <strong>of</strong> Cobitis Linnaeus, 1758: 303; type species:<br />

Cobitis bilseli Battalgil, 1942: 292, by original designation).<br />

Gender feminine.<br />

Taxonomic notes. Cobitis is obviously a polyphyletic genus.<br />

Several chromosome and molecular studies (e.g. Šlechtová<br />

et al., 2008; Kitagawa et al., 2011; Perdices et al., 2012)<br />

show a very complex situation and future research will certainly<br />

result in important changes in the taxonomy <strong>of</strong> Cobitis<br />

and the related genera (Iksookimia, Niwaella, Kichulchoia, etc.).<br />

Nomina nuda<br />

Cobitis longimanus Dybowski, in Sinicyn, 1900: 49 (nomen<br />

nudum; material: “Imperial Warsaw University” 2773;<br />

locality: Onon River)<br />

Cobitis poecilopleura Bleeker, 1872: 146 (nomen nudum;<br />

locality: China)<br />

Nomen nudum<br />

Cobitis yeni Nguyen, 1982: 26 (nomen nudum)<br />

Cobitis yeni Nguyen (T. T.), in Nguyen (V. H.), 2005: 227,<br />

fig. 114 (not available; locality: Vietnam: Ha Tinh Province:<br />

stream Ngam Pho at Huong Son)<br />

Nomenclatural notes. Nguyen (V. H.; 2005: 227) listed<br />

Nguyen (T. T., 1983: 83, pl. 3 fig. 2) as author <strong>of</strong> C. yeni.<br />

Nguyen (1983) is an unpublished thesis and the name is not<br />

available from it. As the description in Nguyen (2005) is<br />

from Nguyen (1983), I treat the author <strong>of</strong> the nomen nudum<br />

as Nguyen (T. T.), in Nguyen (V. H.). The Code art. 16.1<br />

requires that, after 1999, a new name must be explicitly indicated<br />

as intentionally new. This is not the case for C. yeni<br />

in Nguyen (2005) and the name is not available. Further, to<br />

be available, a new specific name published after 1999 must<br />

be accompanied by the explicit designation <strong>of</strong> a holotype or<br />

syntypes (Code art. 16.4). Nguyen (2005) mentioned that<br />

the description is based on 19 specimens but they are not<br />

mentioned as a holotype or syntypes.<br />

Species inquirenda<br />

4.5.1 Cobitis lachnostoma (Rutter, 1897)<br />

Acanthopsis lachnostoma Rutter, 1897: 60 (type locality:<br />

China: Guangdong: Swatow; holotype: CAS-SU 1812,<br />

Böhlke, 1953: 39; compound noun, indeclinable)

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