Conspectus cobitidum - Raffles Museum of Biodiversity Research

10.4.8 Barbatula ‘nuda’, China: Jilin: Antu. Two syntopic species.
Above, with numerous papillae on body (ASIZB 132651,
75.4 mm SL, possibly real B. nuda); below, with smooth body
(ASIZB 132648, 101.6 mm SL; topotype of B. kirinensis). See
also differences in nostrils and respective positions of dorsal and
pelvic fins.
10.4.8 Barbatula nuda, MNHN 1450, 115 mm TL, holotype; China:
Hebei: Xiwanzi.
10.4.12 Barbatula sawadai (Prokofiev, 2007)
Orthrias sawadai Prokofiev, 2007a: 84, fig. 26 (type locality:
Mongolia: Yenisei River drainage: Lake Baikal basin:
Selenge River system: Ero River catchment: stream
Uram; holotype: ZISP 52054; noun in genitive, indeclinable)
10.4.13 Barbatula sturanyi (Steindachner, 1892)
Nemachilus Sturanyi Steindachner, 1892a: 131 (type locality:
FYROM: Lake Ohrid at Pestani, between Ohrid City
and Sveti Naum Monastery; also in Steindachner, 1892b:
378, pl. 2 fig. 3; holotype: NMW 48440; noun in genitive,
indeclinable)
10.4.14 Barbatula tomiana (Ruzsky, 1920)
? Nemacheilus sibiricus Gratzianow, 1907: 167, 168 (type
locality: Russia: Altay: Bija [Biya] River near Bijsk
[Biysk; 52°31'30"N 85°13'30'E]; syntypes: ZMMU
P-2532 [3], Vasil'eva et al., in Pavlinov & Borissenko,
2001: 25; adjective, -us, -a, -um)
Nemacheilus barbatulus tomianus Ruzsky, 1920: 36, figs.
1–2 (type locality: Russia: Siberia: Tomsk oblast: Tom
River at Tomsk, Kuznetsk and Kondoma / brook Kanzas,
a right-hand tributary of Balyk-Su, upper reaches of
Tom River / Ushayka River / Kislovka River near
Tomsk / upper Torsya / upper reaches of Ob itself in area
of Altay (rivers Katun, Cherga, Ursul, Charysh) / in Yenissei
(rivers Abakan and Minusinka); syntypes: LU; adjective,
-us, -a, -um)
? Barbatula barbatula toni morpha tigris Gundriser, 1975:
24 (infrasubspecific, name not available; locality: Russia:
Tuva: lake and bassin of Ii-Kem River [maybe Khemchik,
in Yenisei drainage; 51°16'40"N 90°47'00"E ?])
Taxonomic notes. See under B. toni.
10.4.15 Barbatula toni (Dybowski, 1869)
Cobitis Toni Dybowski, 1869: 957, pl. 18 fig. 10 (type locality:
"common in both river systems" [Russia: Zabaykal-
Kottelat: Conspectus cobitidum
80
10.4.14 Barbatula tomiana, CMK 22367, 118.2 mm SL; Russia:
Altai: Ob drainage: Sema.
10.4.15 Barbatula toni, CMK 19616, 79.9 mm SL; Mongolia:
Amur drainage: Onon.
sky Krai [Transbaikal]: Onon and Ingoda Rivers, Amur
River drainage]; syntypes: ZISP ?, MNHN B.738 [8],
B.739 [4], NMW 50473 [2], Bertin & Estève, 1948: 98,
Eschmeyer & Fricke, 2010; noun in genitive, indeclinable)
Taxonomic notes. Obviously a heterogeneous assemblage
of several species. Considered as a subspecies of B. barbatula
by Prokofiev (2003a, 2007a) but the scale of analysis is
not detailed enough to be informative. Prokofiev (2007a)
recognised a subspecies B. b. toni extending from Japan to
the Urals, with a broad range of morphological variation. I
identify as B. toni the species that seems to be restricted to
the upper Amur drainage (the type locality), recognised by
the backward position of the dorsal fin, the sharply decreasing
body depth behind the dorsal-fin base, and the distinctive
colour pattern with very irregular bars across the back,
on a yellowish background (see, e.g. Bogutskaya et al., 2008:
339; Kottelat, 2006: 99). I have examined B. 'toni' only from
a few places of its range and I have not yet a complete overview
of its variability. But my observations on populations
from Mongolia, Korea and northern China show that the
Selenge (Yenisei drainage) has a distinct species (figured as
"B. b. t. infrasubsp. tomianus" by Berg, 1933: 552, 1948:
871). I tentatively retain the name B. tomiana for it, but
N. sibiricus might be an older available name. The B. 'toni'
from the upper Irtysh (Bulgan River) in Mongolia is distinct
and I tentatively retain the name B. markakulensis for it; this
remains to be confirmed by comparison with fresh material
from Markakul Lake. The B. toni from the eastern slope of
Korea (e.g. Kim & Park, 2007: 193) is apparently the same
as the unnamed species figured by Bogutskaya et al. (2008:
339) and possibly conspecific with populations from Hokkaido,
then B. oreas is the oldest name availabe for it. See
under B. nuda for populations from the western slope of
Korea and northern China.
10.4.16 Barbatula zetensis (Soric, 2001)
Orthrias barbatulus zetensis Šorić, 2001: 66, fig. 1 (type
locality: Montenegro: Vis brook near Vis village, 5–6 km
west of Danilovgrad, tributary of Zeta River; holotype:
BIKU; adjective, -is, -is, -e)
10.5 Claea Kottelat, 2011
Oreias Sauvage, 1874: 334 (type species: Oreias dabryi Sau-