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Conspectus cobitidum - Raffles Museum of Biodiversity Research

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10.3.1 Afronemacheilus abyssinicus, 39.7 mm SL; Ethiopia: Lake<br />

Tana: Blue Nile source. (Photograph by Alexander Golubtsov).<br />

10.4 Barbatula Linck, 1790<br />

Barbatula Linck, 1790: 38 (type species: Cobitis barbatula<br />

Linnaeus, 1758: 303, by absolute tautonymy). Gender<br />

feminine.<br />

Cobites Swainson, 1839: 190, 310 (type species: Cobitis<br />

barbatula Linnaeus, 1758: 303, by monotypy). Gender<br />

feminine.<br />

Orthrias Jordan & Fowler, 1903: 769 (type species: Orthrias<br />

oreas Jordan & Fowler, 1903: 769, by original designation).<br />

Gender masculine.<br />

Nomenclatural notes. Bis repetita placent. Linck (1790)<br />

proposed a classification <strong>of</strong> fishes based on teeth. He split<br />

the Cobitis <strong>of</strong> previous authors into two genera: Cobitis and<br />

Barbatula. Linck used the binominal system and Barbatula<br />

is clearly intended as a generic name. Linck included a single<br />

species in Cobitis (C. heteroclita, p. 33) and two species<br />

in Barbatula (C. barbatula, C. taenia). Although there is no<br />

generic diagnosis, the name is available by indication [Code<br />

art. 12.2.5] as two available names are included. Linck wrote:<br />

"Barbatula. Hierher gehören Cobitis Barbatula Taenia"<br />

[Barbatula. Here belong [plural] Cobitis Barbatula Taenia].<br />

Some have argued that "Cobitis Barbatula Taenia" means<br />

that only one species was included, that this species is<br />

C. taenia and that C. taenia is type species <strong>of</strong> Barbatula.<br />

This is erroneous. Linck used "gehören" [belong], which is<br />

a plural and means that the subject <strong>of</strong> the verb is a plural,<br />

thus that he placed more than one species in Barbatula, and<br />

that these species are barbatula and taenia. If he had included<br />

a single species he would have used the singular "gehört"<br />

[belongs]. Therefore C. barbatula is type species <strong>of</strong> Barbatula<br />

by absolute tautonymy.<br />

Today, "Cobitis Barbatula Taenia" may appear as a<br />

strange combination but not in the context <strong>of</strong> the late 18 th<br />

century, when there were no nomenclature rules, when writing<br />

was less formalised, and when typographical or typesetting<br />

errors were not rare.<br />

Some authors have attributed the authorship <strong>of</strong> Barbatula<br />

to Jordan (1917: 49). As seen from the above, this is not<br />

correct. In addition, a 'Barbatula Jordan, 1917' would be a<br />

junior homonym <strong>of</strong> Barbatula Lesson, 1837: 292 (in Aves)<br />

and is not available (Some authors refer to Lesson, 1831:<br />

164 but Barbatula is not used in that work, only the vernacular<br />

name 'barbion').<br />

10.4.1 Barbatula altayensis Zhu, 1992<br />

Barbatula altayensis Zhu, 1992: 241, figs. 1–2 (type locality:<br />

China: Xinjiang: Kelang He, tributary <strong>of</strong> Ertix He [Irtysh],<br />

near Altay City, 47°52'N 88°06'E; holotype: NGI<br />

888006; adjective, -is, -is, -e)<br />

THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />

77<br />

10.4.2 Barbatula barbatula, CMK 16892, 56.8 mm SL; Switzerland:<br />

Jura: Rhône drainage.<br />

10.4.13 Barbatula sturanyi, CMK 19463, 45.4 mm SL.<br />

10.4.2 Barbatula barbatula (Linnaeus, 1758)<br />

Cobitis Barbatula Linnaeus, 1758: 303 (based on specimens<br />

and Artedi [1738: gen. 2 [4], syn. 2, Cobitis tota glabra<br />

...] and Linnaeus [1746: 125, n. 332, idem]; type locality:<br />

Europae, Asiae, aquis dulcibus; syntypes: NRM 68 [3],<br />

Fernholm & Wheeler, 1983: 282 [listed as non-types],<br />

Lundberg & Svanberg, 2010: 157, fig. 4; noun in apposition,<br />

indeclinable [species names with capitalized initial<br />

in Linnaeus (1758) are nouns])<br />

Cobitis Barbatula Parisiensis La Pylaie, 1835: 534 (nomen<br />

nudum; locality: France: Paris)<br />

Cobitis Barbatula Pictava La Pylaie, 1835: 534 (nomen<br />

nudum; locality: France)<br />

Cobitis furstenbergii Bonaparte, 1846: 26 (nomen nudum;<br />

locality: eastern Germany [probably now Poland])<br />

Cobitis Fürstenbergii Heckel & Kner, 1858: 301 (cited in<br />

synonymy, name not available; author indicated as "Fitzinger<br />

(Prodr. Faun. Austr.)", which apparently is Fitzinger<br />

[1832], but the name does not appear in this work)<br />

Cobitis pironae Nardo, 1866: 143 (type locality: Italy: Basso<br />

Friuli; type material: NT ?; noun in genitive, indeclinable)<br />

Cobitis variabilis Günther, 1868: 355 (on a label in collection,<br />

listed in synonymy, name not available; locality:<br />

Czechia: Moravia)<br />

Nemachilus barbatulus var. taurica Kessler, 1877: 171, 271<br />

(type locality: Ukraine: Crimea: Salgir River; syntypes:<br />

ZISP 3029 [3], 3032 [3], Eschmeyer & Fricke, 2010;<br />

adjective, -us, -a, -um)<br />

? Nemachilus barbatulus caucasicus Berg, 1899: 26, 67 (type<br />

locality: Russia: Chechnya: Argun River, a tributary <strong>of</strong><br />

Terek; types: LU; adjective, -us, -a, -um)

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