Conspectus cobitidum - Raffles Museum of Biodiversity Research

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10.46.22 Yunnanilus pleurotaenia, CMK 13100, 45.5 mm SL; China: Yunnan: Lake Dianchi basin. 10.46.21 Yunnanilus parvus Kottelat & Chu, 1988 Yunnanilus parvus Kottelat & Chu, 1988a: 77, fig. 13 (type locality: China: Yunnan: Kaiyuan County: Nan Tong cave, 23°39'N 103°17'E; holotype: KIZ 847174; adjective, -us, -a, -um) 10.46.22 Yunnanilus pleurotaenia (Regan, 1904) Nemachilus pleurotaenia Regan, 1904: 192 (type locality: China: Yunnan: Sea of Tien at Yunnan Fu [Lake Dianchi, Kunming]; lectotype: BMNH 1904.1.26.36, designated by Kottelat & Chu, 1988a: 82; compound noun, indeclinable) ? Yunnanilus tigeriveinus Li & Duan, 1999: 254, fig. 1 (type locality: China: Yunnan: suburbs of Kunming, in an "opening of underground water", 25°10'N 102°55'E; holotype: HRAS 9901004; also spelt tigerivinus p. 256, as tigeriveinus is used five times and tigerivinus once, tigerivinus is treated as an inadvertent error, thus incorrect original spelling [Code art. 32.5.1]; etymology unknown, treated as noun in apposition, indeclinable) Kottelat: Conspectus cobitidum 136 10.46.23 Yunnanilus pulcherrimus Yang, Chen & Lan, 2004 Yunnanilus pulcherrimus Yang, Chen & Lan, 2004: 112, fig. 1 (type locality: China: Guangxi: Du'an County: Hongshuihe River, a tributary of Xijiang [23°56'N 108°05'E; Romero et al., 2009: 274]; holotype: KIZ 995001; also in Lan, 2004; adjective, -us, -a, -um) Yunnanilus zebrinus Lan & Zhang, in Zhou & Zhang, 2006: 87 (nomen nudum; bibliographic reference make it clear that it is an inadvertent error for Yunnanilus pulcherrimus Yang, Chen & Lan, 2004: 112; fig. II-19 is fig. 1 of Yang et al., 2004: 112) Taxonomic notes. Possibly a species of Micronemacheilus. 10.46.24 Yunnanilus sichuanensis Ding, 1995 Yunnanilus sichuanensis Ding, 1995: 253, fig. 1 (type locality: China: Sichuan: Mianning County: Anning River, Shuyalong Jiang River, 2020 masl, 28°41'N 102°12'E; holotype: MSINR 930026; adjective, -is, -is, -e) 10.46.25 Yunnanilus spanisbripes An, Liu & Li, 2009 Yunnanilus spanisbripes An, Liu & Li, 2009: 631, fig. 1 (type locality: China: Yunnan: Zhanyi County: Niulanjiang River, 25°59'N 103°36'E [Yangtze drainage]; holotype: HRAS 9206064; also spelt spanitripes, p. 637; as first reviser I select spanisbripes as correct original spelling; etymology unknown, noun in apposition, indeclinable)
THE RAFFLES BULLETIN OF ZOOLOGY 2012 APPENDIX: NEW GENUS-GROUP AND FAMILY-GROUP NAMES Ambastaia, new genus Type species. — Botia nigrolineata Kottelat & Chu, 1987. Diagnosis. — Ambastaia is distinguished from all other genera of the family Botiidae by its unique colour pattern in adults consisting in a whitish-yellowish background with a black midlateral stripe on the flank and another, middorsal stripe, with vertical bars connecting the two stripes and extending on the lower half of the body. In A. nigrolineata only the two stripes are present in juveniles and the bars appear at about 40–50 mm SL. In A. sidthimunki, the mid-dorsal stripe is often longitudinally divided by a whitish median area, or a row of whitish blotches, and the midlateral stripe may appear as a row of closely connected blotches; the juveniles have not yet been described. Etymology. — Named for Ambastai (or Ambastus in Latin), a river in Ptolemy’s (ca. 90–168) Γεωγραφκ γφγησις (Geographikê Hyphêgêsis, Geography). This river has been identified as the Mekong (van der Meulen, 1974, 1975). Gender feminine. Remarks. — The colour pattern of A. nigrolineata and A. sidthimunki and its ontogeny are unique within Botiidae. Both were originally described in the genus Botia and they were later transferred to Yasuhikotakia by Nalbant (2002). Still the colour pattern distinguishes them from that of all species of Yasuhikotakia, which includes a large blackish blotch at the base of the caudal fin and, in juveniles, a number of narrow bars on the flank (see figures in Kottelat, 2001b: 87). In most individuals, these bars become indistinct with age. Other elements of the colour pattern (e.g. middorsal stripes, black spots and subdistal margin on fins) may be present. This contrasts with Ambastaia in which the stripes appear first, followed by the vertical elements. Those species of Yasuhikotakia that have been studied osteologically have a long process of the frontal along the anterior margin of the orbit (called “spinous fringe” by Taki, 1972); it is missing in A. sidthimunki. Based on this and other characters, Taki (1972: 78) commented that A. sidthimunki “appears to be the remotest from [...] the rest in the group” [his modesta group, equivalent to Yasuhikotakia]. This condition of the ‘spinous fringe’ has not yet been checked in all known species and therefore it is not used here as a diagnostic character. The molecular phylogeny presented by Šlechtová et al. (2005) included A. nigrolineata and A. sidthimunki and showed that they are each other’s closest relative but not closely related to the other species placed in Yasuhikotakia. On the contrary, they form the sister-group of Sinibotia and together they are the sister-group of Syncrossus. This lineage, in turn, constitutes the sister group of Yasuhikotakia. This clearly 137 shows that A. sidthimunki and A. nigrolineata form a distinct lineage. With the recognition of Ambastaia, Yasuhikotakia seems now monophyletic. Admittedly, it is quite weak to diagnose a genus by its colour pattern; combined with the molecular evidence it seems however justified to formally recognise this lineage by a name awaiting for (possible) more detailed anatomical studies. Assuming that this kind of research will still get support and be tolerated. I wish to point that to me, in botiids, the ontogeny of the colour pattern seems much more informative than the colour pattern itself. Unfortunately there is no published information on the colour pattern of the juveniles of the Chinese genera (Sinibotia, Leptobotia, Parabotia). Theriodes, new genus Type species. — Acanthophthalmus sandakanensis Inger & Chin, 1962: 120. Diagnosis. — Theriodes is distinguished from the other genera of the family Cobitidae in Southeast Asia in having the black marking at the base of the caudal-fin made of a single small black spot; it is slightly vertically elongated, at midheight of the caudal-fin base. The gill opening is small, oval, its length is about equal to the length of the base of the pectoral fin and it is located entirely above the base of the pectoral fin. The anterior nostril is at the tip of a short conical tube and the posterior nostril is not immediately adjacent to the tube, but separated by a distance about equal to its own diameter (shared with Kottelatlimia katik). The pectoral fin has 5 branched rays. In the male the first two pectoral rays are thickened and have a few tubercles along the anterior edge; the second ray is branched but there is no membrane between the branches and the posterior branch is much thinner than the anterior one; superficially the ray may appear unbranched (as originally described by Inger & Chin, 1962); the branches are more discernible in females and juveniles. The other pectoral rays are clearly branched, but the branching point is close to the tip of the rays. The following characters also help to distinguish the genus, although none is unique to it: pelvic fin with 1 simple and 4 branched rays; dorsal-fin origin about halfway between pelvic-fin origin and anal-fin origin; caudal fin rounded; lower lip interrupted medially, each half with an inner, thickened mental lobe; membrane connecting mental lobe to corner of mouth with two barbel-like projections, one of them continuing the mental lobe; barbels short, broad, somewhat triangular and flattened; bifid suborbital spine; eye covered by skin; 25–26 + 10–11 = 35–36 vertebrae (Kottelat & Lim, 1992: 202).
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THE RAFFLES BULLETIN OF ZOOLOGY 201
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Loaches, fishes of the suborder Cob
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others may disagree and might prefe
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priate but, in order to simplify te
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others Grigorii and Vladimir Grumm-
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name available is called the origin
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MCSNC Museo Civico di Storia Natura
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1.1.1 Gyrinocheilus aymonieri, CMK
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2.2.5 Botia kubotai, MHNG 2664.24,
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China: Guangxi: Guilin: syntypes: I
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3.1.1 Vaillantella cinnamomea, CMK
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leptes on p. 311, first reviser [Na
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and C. biwae has been used in a lar
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4.5.54 Cobitis strumicae, CMK 17331
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4.5.51 Cobitis splendens Erk'akan,
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4.9.2 Kottelatlimia katik, ZRC unca
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4.10.5 Lepidocephalichthys berdmore
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4.13 Misgurnus La Cepède, 1803 Mis
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4.14.1 Neoeucirrhichthys maydelli,
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4.16.18 Pangio longimanus Britz & K
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ures copied from earlier works and
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Family ELLOPOSTOMATIDAE 5 Elloposto
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Nomenclatural notes. Silas (1953: 2
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7.3.1 Bhavania australis, CMK 9312,
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7.5.1 Hemimyzon confluens, CMK 2271
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(type locality: India: Manipur: Chi
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7.9.3 Jinshaia sinensis, CMK 13067,
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locality: Vietnam: Lai Chau Provinc
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8 Gastromyzontidae Fowler, 1905 Gas
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8.3.1 'Erromyzon' pachychilus, KIZ
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it should have been in the singular
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River; holotype: FMNH [ex 108270];
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8.7.2 Hypergastromyzon humilis, CMK
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8.11.3 Paraprotomyzon multifasciatu
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ity: China: western Fukien [Fujian]
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Parasewellia polylobata Nguyen [H.
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? Vanmanenia trifasciodorsalata Ngu
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anus is a real difference but partl
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10.3.1 Afronemacheilus abyssinicus,
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42 miles east of Paotow [Baotou, 40
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10.5.1 Claea dabryi, CMK 13064, 67.
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China: Xinjiang: Yarkand River drai
Page 85 and 86: 10.12.1 Ilamnemacheilus longipinnis
Page 87 and 88: Kessler, 1874: 59, by original desi
Page 89 and 90: 10.19.1 Metaschistura cristata, FSJ
Page 91 and 92: 10.21.9 Nemacheilus fasciatus (Vale
Page 93 and 94: 10.22.1 Nemachilichthys ruppelli, C
Page 95 and 96: 10.24.11 Oreonectes retrodorsalis L
Page 97 and 98: 10.25.8 Oxynoemacheilus bureschi, C
Page 99 and 100: 29°30'E; holotype: HUIC MAR-7g; ad
Page 101 and 102: 10.28.2 Paraschistura bampurensis,
Page 103 and 104: 10.30.4 Physoschistura pseudobrunne
Page 105 and 106: 10.32.2 Pteronemacheilus meridional
Page 107 and 108: 10.34.29 Schistura callidora Bohlen
Page 109 and 110: 10.34.76 Schistura isostigma, CMK 1
Page 111 and 112: 10.34.80 Schistura kaysonei Vidthay
Page 113 and 114: 10.34.104 Schistura manipurensis (C
Page 115 and 116: 10.34.144 Schistura punctifasciata,
Page 117 and 118: 10.34.155 Schistura rupecula McClel
Page 119 and 120: 10.35.1 Sectoria atriceps, CMK 1604
Page 121 and 122: 10.36 Seminemacheilus Banarescu & N
Page 123 and 124: 10.41.2 Traccatichthys taeniatus, C
Page 125 and 126: 116°40'E]; lectotype: ZISP 2474, d
Page 127 and 128: or of inadvertence, thus incorrect
Page 129 and 130: 10.42.72 Triplophysa nasobarbatula
Page 131 and 132: locality: Kazakhstan: Alma-Atinskay
Page 133 and 134: 10.44.1 Tuberoschistura baenzigeri,
Page 135: 10.46.7 Yunnanilus discoloris Zhou
Page 139 and 140: Material. — Theriodes sandakanens
Page 141 and 142: tová et al. (2007) and Bohlen & Š
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Page 145 and 146: and adjacent countries]. Izdatelstv
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Page 149 and 150: Zoological Society of London, 1870(
Page 151 and 152: in the Swedish Museum of Natural Hi
Page 153 and 154: logiceskago Muzeja Imperatorskoj Ak
Page 155 and 156: designated as the type species, and
Page 157 and 158: M. Kashiwagi & T. Okazaki, 2005. Ge
Page 159 and 160: Journal of the Bombay Natural Histo
Page 161 and 162: Mai, D. Y., 1978. [Identification o
Page 163 and 164: Ng, H. H. & M. Kottelat, 1999. Oreo
Page 165 and 166: Chinese provinces. Bulletin of the
Page 167 and 168: Roberts, T. R., 1982. The Bornean g
Page 169 and 170: enachbarten Länderstriche. Verhand
Page 171 and 172: skogo Gosudarstvennogo Universiteta
Page 173 and 174: Whitehead, P. J. P., 1971. Correct
Page 175 and 176: (Teleostei: Balitoridae). Ichthyolo
Page 177 and 178: angeli, Triplophysa 10.42.6 anglica
Page 179 and 180: unneanus, Nemacheilus 10.30 brunnea
Page 181 and 182: costata, Diplophysa 10.17.1 costata
Page 183 and 184: Gastromyzon venustus 8.5.34 — vir
Page 185 and 186: japonica, Cobitis 4.5.8 jarutanini,
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macrositanus, Yunnanilus 10.46.14 m
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Nemacheilus dunckeri 10.21.25 — e
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Oreias 10.5 Oreias crassipedunculat
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polylepis, Beaufortia 8.2.13 polyle
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Schistura menanensis 10.34.106 —
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Tarimichthys edsinicus 10.40.2 —
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yaoshanensis, Formosania 8.17.11 ya
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Conspectus cobitidum - Raffles Museum of Biodiversity Research

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