the cynipoid genus paramblynotus - American Museum of Natural ...

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104 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 304 cence. Anterior tentorial pits distinct. Clypeus punctate. Lateral occipital carina not reaching to posterior part of vertex. Occiput glabrous. Anterior flange of pronotum transversely striate; submedian pronotal depressions separated medially. Anterior plate of pronotum glabrous mostly and punctate dorsomedially. Pronotum dorsomedially not raised, distinctly lower than mesoscutum; pronotal crest not raised into a median process. Lateral pronotal carina distinct, reaching pronotal crest dorsomedially. Lateral surface of pronotum foveate-reticulate with pubescence and distinctly separated in the middle. Dorsal pronotal area glabrate, anteriorly with indistinct transverse striation, visible to end of posterior margin of pronotum. Mesoscutum strongly arched dorsally and heavily foveatereticulate. Scutellar sulcus divided by a median longitudinal carina. Mesoscutellum foveate-reticulate; laterodorsal process of mesoscutellum not prominent; posterior margin projected and truncate in dorsal view. Pubescence in axillar area distinct but not conspicuous. Mesopleural triangle densely pubescent and well defined ventrally by a smoothly curved carina. Median mesopleural impression percurrent with a few indistinct transverse costae; mesopleuron glabrous; lower mesopleuron strongly depressed along ventral margin and sparsely pubescent ventrally. Metepisternum areolatereticulate in upper part and conspicuously pubescent ventrally. Lateral propodeal carina percurrent, dorsomedially strongly raised and distinctly curved laterad; median propodeal area glabrate with sparse pubescence; median longitudinal carina purcurrent and submedian transverse carina prominent. Rs+M of forewing arising from middle of basal vein. Marginal cell 3.1 times as long as wide and slightly longer than submarginal cell. Bulla on Sc+R 1 absent. Petiole as long as wide in lateral view. Tergum 8 entirely covered by T7; relative length of T3–7: 1.4:1.0:1.2:1.9:1.5; T3–5 glabrous; T6 finely punctate; T7 finely punctate, with row of sparse pubescence anteriorly. Tibiae and tarsi conspicuously pubescent. Apical teeth of metatibia long, slender, and pointed apically. 1mt/2–5mt 5 0.70. MALE: Length 2.8 mm. Antenna 14-segmented. Pale ring of antenna absent. Paramblynotus filippae is very similar to P. yuani and P. stigi, and can be distinguished from both by several characters as discussed under P. yuani. TYPE MATERIAL: HOLOTYPE: R, Indonesia: Sumatra, Aceh Gunung Leuser National Park, Ketambe Research Station (3u419N, 97u399E, 300 m, Rain Forest, Young Forest Terrace 3, light gap), 1990-II-1–28, D.C. Darling coll. (ROM). PARATYPES: 3RR, 1=: 1R, data as holotype 1991-VI-17–29 (ROM); 2RR, data as holotype, 1991-VI-15–VIII-15 (ROM); 1=, Malaysia: Sabah, near Danum Valley Field Center (ca. 140 m), 1987-II-24– III-24, C. v. Achterberg coll. (RMNH). DISTRIBUTION: Indonesia: Sumatra. ETYMOLOGY: This species is named after Filippa Ronquist, daughter of F.R. Paramblynotus shimenensis, new species FEMALE: Length 4.5 mm. Body entirely dark brown. Submedial pale ring of antenna consisting of F6–7, F8–11 almost black, and all other segments dark brown. Forewing with a deep ferruginous macula covering marginal cell and the part of third cubital cell behind marginal cell. Body parts, except as specified below, with moderately dense silvery pubescence. Antenna 13-segmented; flagellum filiform. Vertex foveate-reticulate. Eye prominent and distinctly extended laterally beyond outer margin of gena. Ocellar plate slightly raised, glabrate anteriorly, foveate posterior to anterior ocellus, and with a row of indistinctly defined foveae along interior side. Median frontal carina complete to clypeus and raised into a distinct lamilate process between antennal sockets. Upper face foveate-rugose; antennal scrobe defined by distinct carina laterally, distinctly depressed, and glabrate and finely punctate with apparent fine longitudinal striation. Gena areolate/foveate-reticulate with a median vertical carina. Lower face and clypeus foveate-rugose. Anterior tentorial pits indistinct. Lateral occipital carina reaching posterior part of vertex. Occiput glabrous. Anterior flange of pronotum transversely striate; submedian pronotal depressions sep-
2007 LIU ET AL.: REVISION OF PARAMBLYNOTUS (HYMENOPTERA) 105 arated medially. Anterior plate of pronotum glabrous, finely sparsely punctate dorsomedially. Pronotum dorsomedially not raised, distinctly lower than mesoscutum; pronotal crest not raised into a median process. Lateral pronotal carinae distinct and reaching pronotal crest dorsomedially; lateral surface of pronotum distinctly separated in the middle. Lateral surface of pronotum foveate-reticulate. Dorsal pronotal area glabrate, present to anterior third of posterior margin of pronotum. Mesoscutum strongly arched dorsally and heavily foveate-reticulate. Scutellar sulcus divided by a median longitudinal carina. Mesoscutellum foveatereticulate; laterodorsal process of mesoscutellum smoothly curved lateroposteriorly to posterior margin of mesoscutellum; posterior margin broadly rounded in dorsal view. Pubescence in axillar area distinct but not conspicuous. Mesopleural triangle densely pubescent, well defined ventrally by a smoothly curved carina. Median mesopleural impression percurrent, with two more or less distinct transverse costae; mesopleuron glabrous; lower mesopleuron strongly depressed along ventral margin, and densely pubescent ventrally. Metepisternum areolate-reticulate in upper part and conspicuously pubescent ventrally. Lateral propodeal carina percurrent, distinctly raised dorsally throughout and slightly curved laterad in posterior half; median propodeal area glabrate with sparse pubescence; median longitudinal carina present anteriorly only, submedian transverse carina prominent, and two submedian longitudinal carinae present posteriorly. Rs+M of forewing arising from middle of basal vein. Marginal cell 2.7 times as long as wide and slightly longer than submarginal cell. Bulla on Sc + R 1 absent. Petiole 1.1 times as long as wide in lateral view. Tergum 8 completely hidden beneath T7; relative length of T3–7: 2.0:1.0:1.2:2.5:1.2; T3–5 glabrous; T6 finely punctate with single row of pubescence; T7 finely punctate, with a band of sparse pubescence anteriorly. Tibiae and tarsi conspicuously pubescent. Apical teeth of metatibia long, slender, and pointed apically. 1mt/ 2–5mt 5 0.56. MALE: Unknown. Paramblynotus shimenensis differs from all other species of annulicornis species complex by combination of the following characters: (1) median frontal carina raised into a distinct lamilate process between antennal sockets and complete to clypeus, (2) gena areolate/ foveate-reticulate with a median vertical carina, and (3) T6 longer than T3. TYPE MATERIAL: HOLOTYPE: R, China: Hunan, Shimen, Jiangping, Xiaoxi (1,470 m, mixed evergreen and deciduous broad-leaved forest), 1992-VIII-16–IX-15, F. Du and Z. Liu coll. (ZICA). DISTRIBUTION: China: Hunan. ETYMOLOGY: The species is named after the type locality. Paramblynotus glaberus, new species FEMALE: Length 3.5 mm. Body entirely yellow brown. Submedial pale ring of antenna consisting of F6–8, F9–11 dark brown, and all other segments brown. Forewing with a deep ferruginous macula covering marginal cell and the part of third cubital cell behind marginal cell. Body parts, except as specified below, with medially dense silvery pubescence. Antenna 13-segmented; flagellum filiform. Vertex foveate-reticulate. Eye prominent and distinctly extended laterally beyond outer margin of gena. Ocellar plate slightly raised, defined by distinct carinae laterally, foveate posterior to anterior ocellus, and glabrate anteriorly. Median frontal carina complete to clypeus and raised between antennal sockets to form a distinct lamilate process. Upper face glabrate-foveate; antennal scrobe distinctly depressed, glabrous, and defined by a distinct carina laterally. Gena areolaterugulose with a median vertical carina. Lower face and clypeus foveate-rugose. Anterior tentorial pits indistinct. Lateral occipital carina reaching posterior part of vertex. Occiput glabrous, anteriorly separated by a transverse carina from vertex. Anterior flange of pronotum transversely striate; submedian pronotal pits separated from each other medially. Anterior plate of pronotum glabrous. Pronotum dorsomedially not raised, distinctly lower than mesoscutum; pronotal crest not raised into a median process. Lateral pronotal carina
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THE CYNIPOID GENUS PARAMBLYNOTUS: R
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ABSTRACT The genus Paramblynotus is
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2007 LIU ET AL.: REVISION OF PARAMB
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