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the cynipoid genus paramblynotus - American Museum of Natural ...

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130 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 304<br />

smoothly curved carina delimiting <strong>the</strong> triangle.<br />

Median mesopleural impression percurrent<br />

and glabrate with apparent transverse<br />

carinae. Upper and lower mesopleuron glabrous;<br />

lower mesopleuron also pubescent in<br />

ventral impression. Metepisternum conspicuously<br />

pubescent entirely. Lateral propodeal<br />

carinae percurrent, distinctly divergent posteriorly,<br />

and strongly raised dorsally to form<br />

in <strong>the</strong> middle a distinct triangular process;<br />

median propodeal area conspicuously pubescent<br />

anteriorly and glabrous posteriorly;<br />

median longitudinal carina anterior to <strong>the</strong><br />

median transverse carina barely visible due to<br />

conspicuous pubescence. Rs+M <strong>of</strong> forewing<br />

distinct except slightly nebulous in basal<br />

third, arising from middle <strong>of</strong> basal vein.<br />

Marginal cell 3.5 times as long as wide and<br />

1.7 times as long as submarginal cell. Bulla<br />

on Sc+R 1 absent.<br />

Petiole 0.40 times as long as wide in lateral<br />

view. Tergum 8 completely covered by T7;<br />

relative size <strong>of</strong> abdominal T3–7: 2.3:1.0:1.2.<br />

:3.8:1.4; T3 glabrous; T4–5 glabrous and<br />

finely punctate dorsally; T6 finely and<br />

densely punctate dorsolaterally, with a single<br />

row <strong>of</strong> pubescence in <strong>the</strong> middle; T7 punctate<br />

with pubescence dorsolaterally in anterior<br />

third and glabrous posteriorly. Apical teeth<br />

<strong>of</strong> metatibia slender and pointed apically.<br />

Apical process <strong>of</strong> first metatarsomeres reaching<br />

to two-thirds <strong>of</strong> second metatarsomere.<br />

MALE: Unknown.<br />

Paramblynotus fucosus is unique among all<br />

Paramblynotus species for its very conspicuous<br />

patch <strong>of</strong> silvery hairs covering <strong>the</strong><br />

metepisternum and propodeum, its evenly<br />

ferruginous wings, and its relatively long<br />

apical process <strong>of</strong> first metatarsomeres.<br />

TYPE MATERIAL: HOLOTYPE: R, Papua<br />

New Guinea: Bulolo Dist., 1982, H. Roberts<br />

coll. (NHM).<br />

DISTRIBUTION: Papua New Guinea<br />

ETYMOLOGY: From Latin, fucosus, colored,<br />

painted. The name refers to <strong>the</strong><br />

ferruginous wings <strong>of</strong> <strong>the</strong> species.<br />

ACKNOWLEDGMENTS<br />

This study is an expansion <strong>of</strong> part <strong>of</strong> Z.L.’s<br />

Ph.D. dissertation supported by a scholarship<br />

from <strong>the</strong> Oscar and Lili Lamm’s Memorial<br />

Foundation and was partly supported by<br />

a grant from <strong>the</strong> Swedish <strong>Natural</strong> Science<br />

Research Council to F.R. and G.N. Visits to<br />

insect collections by Z.L. and Meicai Wei in<br />

China were supported by a research grant<br />

from <strong>the</strong> National <strong>Natural</strong> Science Foundation<br />

<strong>of</strong> China to Z.L. (no. 39300107). Part <strong>of</strong><br />

<strong>the</strong> systematic work was conducted when<br />

Z.L. was supported by a Kalbfleisch fellowship<br />

from <strong>the</strong> <strong>American</strong> <strong>Museum</strong> <strong>of</strong> <strong>Natural</strong><br />

History and a Boyd postdoctoral scholarship<br />

at <strong>the</strong> Field <strong>Museum</strong> <strong>of</strong> <strong>Natural</strong> History.<br />

During <strong>the</strong> later revisions <strong>of</strong> <strong>the</strong> manuscript,<br />

Z.L. was funded by <strong>the</strong> Center for Insect<br />

Science through NIH Training Grant 1 K12<br />

Gm00708, EIU Council on Faculty Research<br />

through a 2005 Summer Research Award,<br />

and was supported by <strong>the</strong> Arizona-Sonora<br />

Desert <strong>Museum</strong> in many respects. Z.L. also<br />

thanks David Strait for suggestions during<br />

<strong>the</strong> development <strong>of</strong> <strong>the</strong> FMCK methods and<br />

for making comparisons between <strong>the</strong> FMC<br />

and FMCK methods using real data. The<br />

assistance <strong>of</strong> <strong>the</strong> many curators from various<br />

institutions (listed in material section) with<br />

loans <strong>of</strong> specimens and <strong>the</strong>ir willingness (and<br />

unwillingness) to extend <strong>the</strong>se loans have<br />

greatly helped <strong>the</strong> completion <strong>of</strong> <strong>the</strong> study.<br />

Z.L is grateful to <strong>the</strong> colleagues <strong>of</strong> <strong>the</strong><br />

institutions where he has worked in <strong>the</strong> past<br />

few years, especially former postdoctoral<br />

supervisors Drs. J. Carpenter (AMNH), P.<br />

Goldstein (FMNH), and R. Brusca (ASDM);<br />

this project would be impossible to complete<br />

without <strong>the</strong>ir support and encouragement.<br />

Finally, <strong>the</strong> priviledge <strong>of</strong> Z.L being a Research<br />

Associate <strong>of</strong> <strong>the</strong> <strong>American</strong> <strong>Museum</strong> <strong>of</strong><br />

<strong>Natural</strong> History made <strong>the</strong> publication <strong>of</strong> this<br />

study possible.<br />

REFERENCES<br />

Akaike, H. 1974. A new look at <strong>the</strong> statistical<br />

model identification. IEEE Transactions on<br />

Automatic Control AC-19: 716–723.<br />

Archie, J. 1985. Methods for coding variable<br />

morphological features for numerical analysis.<br />

Systematic Zoology 34: 326–345.<br />

Ashmead, W.H. 1903. Classification <strong>of</strong> <strong>the</strong> gallwasps<br />

and <strong>the</strong> parasitic <strong>cynipoid</strong>s, or <strong>the</strong><br />

superfamily Cynipoidea. IV. Psyche (Cambridge)<br />

10: 210–215.<br />

Axelrod, D.I., and P.H. Raven. 1978. Late<br />

Cretaceous and Tertiary vegetation history <strong>of</strong><br />

Africa. In M.J.A. Werger (editor), Biogeogra-

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