the cynipoid genus paramblynotus - American Museum of Natural ...

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68 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 304 middle legs sparsely punctate with pubescence; hindlegs sparsely punctate with pubescence except metacoxa dorsally glabrous and metatibia and metatarsomeres densely punctate with pubescence. Apical teeth of metatibia long, thin, and pointed apically. 1mt/ 2–5mt 5 0.55. MALE: Unknown. Paramblynotus antistatus can be divided from the othe other species of the immaculatus clade by its distinctly exposed T7 of female metasoma and its smooth gena, except P. scalptus. However, P. antistatus differs from P. scalptus in having vertex and gena glabrous. TYPE MATERIAL: HOLOTYPE: R, Namibia, Windhoek, Regenstein (15 mi SSW), 1972-IV-9 (NHM). DISTRIBUTION: Namibia, Windhoek. ETYMOLOGY: From Latin, anti-, against, and estat, position, standing. This name was coined because of the glabrous vertex and gena of this species, being in distinct contrast to the sculptured vertex and gena of the most similar species, P. scalptus. Paramblynotus scalptus, new species FEMALE: Length 2.8 mm. Head and mesosoma black. Metasoma black with T3 and T4 brown. Antennae yellow with distal three segments dark. Forewing clear in about distal two-fifths, and ferruginous basally. Fore- and middle legs brown to dark brown. Hindlegs dark brown to almost black. Antenna 13-segmented; flagellum gradually expanded apically; median flagellomeres not constricted. Vertex densely punctate medially and diagonally carinate laterally. Eye prominent, distinctly extended laterally beyond outer margin of gena. Distance between posterior ocelli 3.5 times as wide as the distance between posterior ocellus and eye. Ocellar plate not raised and not defined laterally by a carina; median frontal carina only weakly present between antennal sockets. Upper face, including antennal scrobe, coriarious; antennal scrobes only slightly depressed with very slight traces of foveate sculpture, and defined by carina laterally. Lower face foveate-reticulate. Anterior tentorial pits distinct. Clypeo-pleurostomal sulcus and epistomal sulcus almost form an arch. Clypeus distinctly longitudinally carinate. Gena foveate-punctate. Lateral occipital carina indistinct beyond vertex. Occiput glabrous. Anterior flange of pronotum glabrous. Submedian pronotal depressions distinctly separated medially. Anterior plate of pronotum glabrate anteriorly and coriarious posteriorly, laterally also with setigerous punctures. Pronotum dorsomedially not distinctly raised; pronotal crest medially not raised into a process. Lateral pronotal carina distinct, not reaching pronotal crest dorsomedially. Lateral surface of pronotum coarsely foveate-reticulate. Dorsal pronotal area reaching end of posterior margin of pronotum and with closely arranged punctures in a single row. Mesoscutum distinctly arched dorsally and foveate-reticulate. Scutellar sulcus divided by a single median longitudinal carina; mesoscutellum foveate-reticulate; posterior margin narrowly rounded in dorsal view. Mesopleural triangle ventrally well defined by smoothly curved carina and with conspicuous white pubescence. Upper mesopleuron glabrous; speculum longitudinally carinate; and median impression percurrent with multiple unevenly distributed transverse carinae. Lower mesopleuron glabrous, conspicuously pubescent ventrally. Metepisternum impressed and nude above, with an elevated, glabrous-punctate area posteromedially, and somewhat depressed and pubescent ventrally. Propodeum mostly areolate-reticulate; lateral and median propodeal carinae present only anteriorly and about same strength as additional longitudinal carinae posteriorly. Nucha glabrous. Rs+M of forewing nebulous, arising from posterior end of basal vein. Marginal cell 2.0 times as long as wide and slightly longer than submarginal cell. Bulla on Sc+R 1 absent. Abdominal petiole 0.5 times as long as wide in lateral view. Relative length of T3–8: 1.4:1.0:2.0:0.8:0.15:0.3; T3–5 glabrous; T6–7 finely punctate; T7 of metasoma slightly exposed; T8 with a few large setigerous punctures. All legs densely punctate with pubescence except metacoxa dorsally glabrous. Apical teeth of metatibia long, thin, and pointed apically. 1mt/2–5mt 5 0.55.
2007 LIU ET AL.: REVISION OF PARAMBLYNOTUS (HYMENOPTERA) 69 MALE: Length 2.5 mm. Paramblynotus scalptus, as stated above, is similar to P. antistatus, but it can be distinquished by its sculptured vertex and gena. In addition, its forewing is more extensively ferruginous, extending to cover basal three-fifths, instead of basal two-thirds as P. antistatus. TYPE MATERIAL: HOLOTYPE: R, South Africa: Transvaal (Rustenburg Nature Reserve, 25.40S, 27.12E), 1983-XII-8, C.D. Eardley coll. (PPRI). PARATYPES: 1R, 1=: 1R, South Africa, Transvaal, Kruger National Park, Pufuri (22.26S, 31.12E), 1984-I-21, C.D. Eardley coll. (PPRI); 1=, South Africa, East Transvaal (in woodland 15 km east of Klaserie, Guernsey Farm), 1985-XII-19–31, S. and J. Peck coll. (CNCI). DISTRIBUTION: South Africa, East Transvaal. ETYMOLOGY: From Latin, scalpo, cut, carve. The name describes the sculptured vertex and gena of the species as compared to P. antistatus. Paramblynotus vannoorti, new species FEMALE: Length 2.2 mm. Body black except antennae dark yellow. Forewing mostly clear with a wide, medial smoky band covering marginal and submarginal cells and the areas behind; darker in marginal and sumarginal cells and lighter behind. Antenna 13-segmented; flagellum gradually expanded apically; median flagellomeres not constricted. Vertex entirely diagonally carinate. Eye prominent, distinctly extended laterally beyond outer margin of gena. Distance between posterior ocelli nearly 5.0 times as wide as the distance between posterior ocellus and eye. Ocellar plate not raised, defined laterally by a carina, radiating carinate anteriorly and foveate-reticulate posterior to anterior ocellus; median frontal carina extended slightly beyond lower margin of antennal sockets. Upper face, including most of antennal scrobe, coriarious with very sparse fine punctures, and defined by carina laterally. Antennal scrobes shallow, glabrous posteriorly. Lower face coarsely coriarious and superficially foveate-reticulate. Anterior tentorial pits distinct. Clypeo-pleurostomal sulcus and epistomal sulcus almost form an arch. Clypeus glabrous. Gena cariarious. Lateral occipital carina indistinct beyond vertex. Occiput glabrous. Anterior flange of pronotum glabrous. Submedian pronotal depressions fused with each other medially. Anterior plate of pronotum glabrous anteriorly and sparsely, coarsely punctate posteriorly. Pronotum dorsomedially not distinctly raised; pronotal crest medially not raised into a process. Lateral pronotal carina distinct, nearly reaching pronotal crest dorsomedially. Lateral surface of pronotum coarsely foveate-reticulate. Dorsal pronotal area reaching to end of posterior margin of pronotum, glabrous anteriorly, and with closely arranged punctures in a single row. Mesoscutum distinctly arched dorsally and foveate-reticulate. Scutellar sulcus divided by a single median longitudinal carina; mesoscutellum foveatereticulate; posterior margin rounded in dorsal view. Mesopleural triangle ventrally well defined by smoothly curved carina and with conspicuous white pubescence. Upper mesopleuron glabrous; speculum longitudinally carinate; and median longitudinal impression percurrent with multiple unevenly distributed transverse carinae. Lower mesopleuron glabrous, pubescent ventrally. Metepisternum coarsely foveate-reticulate and nude above, with a small elevated, glabrous area medially, and pubescent ventrally. Propodeum mostly areolate-reticulate; lateral and median propodeal carinae present only anteriorly and about same strength as the additional longitudinal carinae present posteriorly. Median propodeal area alveolate anteriorly and longitudinally carinate posterorly. Nucha glabrous. Rs+M of forewing nebulous, arising from close to posterior end of basal vein. Marginal cell 2.0 times as long as wide and slightly longer than submarginal cell. Bulla on Sc+R 1 absent. Abdominal petiole 0.5 times as long as wide in lateral view. Relative length of T3–8: 1.4:1.0:2.0:1.0:0:0.2; T3–4 glabrous; T5 very finely and sparsely punctate, T6–7 finely punctate; T7 completely covered by T6; T8 slightly exposed, with sparse setigerous puctures. All legs sparsely punctate with pubescence except metacoxa dorsally glabrous and metatibia and metatarsomeres densely pu-
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THE CYNIPOID GENUS PARAMBLYNOTUS: R
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ABSTRACT The genus Paramblynotus is
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2007 LIU ET AL.: REVISION OF PARAMB
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