the cynipoid genus paramblynotus - American Museum of Natural ...

2007 LIU ET AL.: REVISION OF PARAMBLYNOTUS (HYMENOPTERA) 67
cence except metacoxa dorsally glabrous and
metatibia and metatarsomeres densely punctate
with pubescence. Apical teeth of metatibia
long, thin, and pointed apically. 1mt/
2–5mt 5 0.64.
MALE: Unknown.
Paramblynotus immaculatus, antistatus,
scalptus, vannoorti, diminutus, angolensis,
and minutus form a very distinct, monophyletic
clade within the trisetosus group.
This clade is characteristic in having (1) very
conspicuous anterior tentorial pits; (2) distance
between posterior ocelli at least three
times as wide as distance between posterior
ocellus and eye; (3) speculum longitudinally
carinate; (4) T5 of female dorsally expanded
and the largest metasomal tergum, T7 largely
or entirely covered by T6, T8 exposed; and
(5) wings with different color patterns; Rs+M
vein of forewing arising from posterior end of
basalis.
Paramblynotus immaculatus can be easily
separated from the other species of this clade
by having (1) wings evenly ferruginous and
without darker bands, (2) mesoscutellum
posteriorly with a broad emargination, and
(3) median frontal carina almost reaching
clypeus.
TYPE MATERIAL: HOLOTYPE: R, Zaire,
Pidigala, 1952-iv-23, H. De Saeger coll.
(NHM).
DISTRIBUTION: Zaire.
ETYMOLOGY: From Latin, im- not, and
macula, mark, spot. The name describes the
evenly ferruginous wings of the species.
Paramblynotus antistatus, new species
FEMALE:
Length 2.1 mm. Body black to
dark brown with antennae and legs yellow to
yellow brown. Forewing brown in about
basal two-thirds and clear beyond the tip of
marginal cell.
Antenna 13-segmented; flagellum gradually
expanded apically; median flagellomeres
not constricted. Vertex largely glabrous with
sparse punctures, slightly rugose laterally.
Eye prominent, distinctly extended laterally
beyond outer margin of gena. Distance
between posterior ocelli 3.0 times as wide as
the distance between posterior ocellus and
eye. Ocellar plate not raised and not defined
laterally by a carina; median frontal carina
extended from anterior ocellus to slightly
beyond lower margin of antennal sockets.
Upper face, including antennal scrobe, coriarious;
antennal scrobes only slightly depressed,
defined by carina laterally. Lower
face coriarious above and transversely rugose
with foveae ventrally. Anterior tentorial pits
distinct. Clypeo-pleurostomal sulcus and
epistomal sulcus almost form an arch.
Clypeus glabrate and weakly longitudinally
carinate. Gena coriarious. Lateral occipital
carina indistinct beyond vertex. Occiput
glabrous.
Anterior flange of pronotum glabrous.
Submedian pronotal depressions distinctly
separated medially. Anterior plate of pronotum
glabrate and coarsely punctate. Pronotum
dorsomedially not distinctly raised;
pronotal crest medially not raised into a process.
Lateral pronotal carina distinct, not
reaching pronotal crest dorsomedially. Lateral
surface of pronotum foveate-reticulate.
Dorsal pronotal area glabrate-punctate and
reaching to end of posterior margin of
pronotum. Mesoscutum distinctly arched
dorsally and foveate-reticulate. Scutellar sulcus
divided by a single median longitudinal
carina; mesoscutellum foveate-reticulate;
posterior margin round in dorsal view.
Mesopleural triangle ventrally well defined
by smoothly curved carina and partly with
white pubescence. Upper mesopleuron glabrous,
distinctly depressed; speculum longitudinally
carinate; median impression percurrent
with three unevenly distributed
transverse carinae. Lower mesopleuron glabrous;
pubescence sparse above and dense
ventrally. Metepisternum foveate-rugose,
nude above and conspicuously pubescent
ventrally. Propodeum, including median propodeal
area, areolate-reticulate; lateral and
median propodeal carinae present only anteriorly;
lateral propodeal carina with distinct
pubescence dorsally. Nucha glabrous. Rs+M
of forewing nebulous, arising from close to
posterior end of basal vein. Marginal cell 2.0
times as long as wide, and as long as
submarginal cell. Bulla on Sc+R 1 absent.
Abdominal petiole 0.3 times as long as
wide in lateral view. Relative length of T3–8:
1.3:1.0:2.0:1.1:0.4:0.4; T3–4 glabrous; T6–7
punctate; T7 of metasoma distinctly exposed;
T8 punctate with a few foveae. Front and