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Crustacea: Copepoda - Cerambycoidea.com

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Epiphyte associates: the algae, mosses, liverworts, lichens and micro-fungi which<br />

use exposed bark surfaces as structure to grow upon provide further difficulties,<br />

mainly through misinterpretation by the human recorders. Plaited door snail<br />

Cochlodina laminata is <strong>com</strong>monly found on tree trunks and branches and may be<br />

found sheltering deep inside wood-decay cavities, but it is most definitely part of the<br />

epiphyte <strong>com</strong>munities not wood-decay. The greatest difficulties occur with predatory<br />

species, as species using the epiphytes as cover and shelter will feed not only on the<br />

inhabitants of the epiphyte cover but also on any wood-decay associates which they<br />

encounter. Many spiders are characteristic of the outer bark of trees and, like plaited<br />

door snail, may shelter deep within decay cavities and even foray for food into the<br />

interior of the tree.<br />

Caches, etc: an interesting note by Whitehead (1986) drew attention to the presence<br />

of skeletal fragments of a wide range of woodland beetles inside a hollowed oak<br />

pollard which were associated with a cache of nuts and seeds gathered by a wood<br />

mouse. This is a useful warning for when attempting to draw conclusions based on<br />

fragments found in wood decay.<br />

Life history and ecology. Decaying wood provides excellent cover or shelter for<br />

many species that do not actually develop within it. Many species which may be<br />

encountered while sampling decaying wood may therefore be irrelevant to this listing,<br />

although the details of their life histories and ecology may be so inadequate that they<br />

are included by mistake. Examples include species that are diurnal and spend the<br />

hours of darkness within wood and vice versa, others that over-winter or aestivate in<br />

deadwood, and even those which pupate in deadwood but develop elsewhere.<br />

Biogeography<br />

There are essentially two major faunas here, the Atlantic version of temperate broad-leaved<br />

forest fauna and the boreal forest fauna. The Atlantic temperate fauna is widespread over<br />

lowland Britain, as far north as southern Scotland; it be<strong>com</strong>es increasingly species poor in the<br />

west, although this is perhaps a reflection of the increasingly "highland conditions" of the<br />

older rocks. In Ireland, the fauna is very much a relict one owing to the very extensive forest<br />

clearances, and the character is something of a hybrid between the temperate and boreal<br />

forests of Britain. The northern parts of Scotland have boreal pine, birch and aspen forests,<br />

with a fauna more akin to that of Scandinavia than to the rest of Britain. A good proportion<br />

of the species are <strong>com</strong>mon to both broad categories, but others are clearly characteristic of<br />

temperate or boreal forest in the British Isles. Relationships with the continental faunas has<br />

not been considered to any great extent.<br />

Structure of the annotated list<br />

The species accounts are a synthesis and summary of a large number of records and<br />

observations, both published and unpublished, and include some ecological analysis of the<br />

available information. It was felt that full citations would make the text unacceptably<br />

cumbersome. The decision was therefore taken not to include details of the source, but to<br />

provide a bibliography of some of the most important sources. The non-aculeate<br />

Hymenoptera is treated as an exception as knowledge of the taxonomy and biology of this<br />

11

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