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laeobotany and Palynology, 2007, 144(3-4):363-3642008040120中 国 南 方 发 现 两 种 工 蕨 新 种 及 用 生 物 地 层 学方 法 测 定 徐 家 冲 组 的 年 代 = Two Species ofZosterophyllum from South China and Dating ofthe Xujiachong Formation with a BiostratigraphicMethod. ( 英 文 ). Wang Deming. ActaGeologica Sinica, 2007, 81(4): 525-533+535-538+534Zosterophyllum longa sp.nov.is reported fromthe Lower Devonian Pingyipu Formation, JiangyouDistrict,northern Sichuan of China.Thisnew plant has vertically elongate sporangia differingfrom all known species of Zosterophyllum,demonstratingmorphological variations inZosterophyllophytina.New materials of Zosterophyllumyunnanicum from the Lower DevonianXujiachong Formation,Qujing District,easternYunnan of China show that fertile axes of thisspecies are anisotomous or pseudomonopodial orK-shaped in bra...2008040121中 国 西 南 地 区 上 二 叠 统 保 存 解 剖 结 构 的 观 音莲 座 蕨 类 : 盘 县 辉 木 ( 新 种 )Psaroniuspanxianensis sp. nov. 的 树 干 化 石 = Anatomicallypreserved marattialean plants from the UpperPermian of southwestern China: the trunk ofPsaronius panxianensis sp. nov.. ( 英 文 ). He X-Y;Wang S-J; Hilton J; Tian B-L; Zhou Y-L. PlantSystematics and Evolution, 2008, 272(1-4): 155-180A new species of the marattialean fern trunkPsaronius Cotta is described from the UpperPermian Cathaysian flora in the Xuanwei Formationof Guizhou Province, SW China, andnamed P. panxianensis sp. nov. Stems possess ahelically diverging sequence of leaf traces organizedin a 2/7 phyllotaxy and are characterizedby stages of leaf trace development that includean early ontogenetic stage with a single vascularbundle (Stewartiopteris-type), a middle stagewith two vascular bundles (Stipitopteris-type),and a late stage with three vascular bundles.Roots diverge singly or occasionally in pairs,arising alternately from the centrifugal surface ofthe peripheral cauline bundles (PCB) near the tip,and form both bound and free root mantles. Thisspecies is distinguished from all previously recognizedspecies from the Euramerican andGondwana floras in having an anchor-shapedsclerenchymatous strand internal to the PCB anda large V-shaped strand formed by the connectionof two anchor-shaped strands. Psaroniuspanxianensis sp. nov. is important as its leaftraces show three distinct ontogenetic developmentstages within the trunk, with this organizationinterpreted as a derived condition withinpsaroniaceous marattialean fern evolution. In thestem abundant air space suggests that P.panxianensis lived in waterlogged substrateswith aerenchyma facilitating gaseous exchange.However, the presence of marattialean ferndominated communities in the Upper Permian ofSW China suggest an overall drying trend withthese plants replacing earlier lycopsid andsphenopsid dominated wetland communities.2008040122华 北 下 二 叠 统 煤 核 中 保 存 解 剖 结 构 的 鳞 木 化石 : 过 渡 无 被 果 孢 ( 新 种 )Achlamydocarpon intermedium sp. nov. =Anatomically preserved lepidodendralean plantsfrom lower permian coal balls of northern China:Achlamydocarpon intermedium sp. nov.. ( 英 文 ).Zhou Yi-Long; Wang Shi-Jun; Hilton J; TianBao-Lin. Plant Systematics and Evolution, 2008,273(1-2): 71-85Lepidodendralean lycopsids, a dominant componentin Late Palaeozoic wetland plant communities,possess a diversity of reproductivestructures that are primarily known from theLate Palaeozoic floras of Europe and NorthAmerica. Here we document an anatomicallypreserved lepidodendralean lycopsid sporophyllwith attached megasporangium from the LowerPermian Taiyuan Formation in Shanxi Province,northern China. The sporophyll has a pedicelonto which the sporangium is attached, and thesporangium is dorsiventrally flattened, proximallydehiscent and longitudinal ridged. Themegasporangial wall comprises three zones: anouter uniseriate layer of columnar cells, a middlelayer 1–3 cells thick comprising isodiametricparenchymatous cells, and an inner zone 1–3cells thick of thick-walled cells. The vascularsystem comprises a single xylem strand surroundedby zone of parenchyma that continuesthrough the pedicel into the lamina. Within themegasporangium a single functional megasporeand three abortive megaspores occur. Features ofthis specimen conform to AchlamydocarponSchumacher-Lambry, and comparisons withother species show it shares similarities with A.takhtajanii (Sni.) Schumacher-Lambry and A.varius Taylor and Brack-Hanes. Although themorphology and anatomy of the specimen wedescribe overlaps with these two species, it isdistinct from both leading to the erection of thenew species A. intermedium sp. nov. The evolutionarysignificance of A. intermedium sp. nov.and the identity of its parent plant are considered,and the status and systematic position of “Orientallepidophytes” from the Cathaysian floras arediscussed.37

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