was pulsed, with major declines usually associatedwith the onset of cold intervals, and partialrecoveries in intervening warm intervals. Thepulses varied in timing between sites. The highestoccurrences (HOs or local disappearances) ofindividual Extinction Group species are variableand mostly diachronous between sites. Therewas a progressively increasing overall rate oflocal disappearances per time through the latePliocene (onset of northern hemisphere glaciation)to the middle Pleistocene climatic revolution,with the peak period of local disappearances(mostly 0.9–0.7 Ma) up to 0.5 Ma earlierat deeper and cooler water locations. The youngestoccurrence of any member of the ExtinctionGroup (Stilostomella extinction datum) is remarkablyconsistent in all sites (0.65–0.57 Ma).The timing of these abundance declines, highestoccurrences (or withdrawals) and extinctionswas essentially the same as in the Atlantic Ocean.The precise mechanistic cause of the Stilostomellaextinction (cooling, increased oxygenationof bottom waters, food supply changes)is yet to be resolved. This study reveals a muchlarger extinction of taxa than previously recorded(middle Pleistocene extinction rate of23% of the bathyal-upper abyssal fauna/myr).Becoming extinct, or virtually so, during thisperiod were at least two families (Stilostomellidae,Pleurostomellidae), one subfamily (Plectofrondiculariinae),at least 17 genera (Awhea,Chrysalogonium, Cribronodosaria, Ellipsoglandulina,Ellipsopleurostomella, Ellipsopolymorphina,Haeuslerella, Mucronina, Myllostomella,Nodosarella, Orthomorphina, Parafrondicularia,Pleurostomella, ?Rectuvigerina, Siphonodosaria,Stilostomella, Strictocostella) and 53 species.Their taxonomy is reviewed, revealing manysynonymies, often going back to Schwager’s(1866) pioneering study of Pliocene deep-seaforaminifera of Car Nicobar, Indian Ocean. Arevised generic subdivision of the Stilostomellidaeis proposed based primarily on aperturalfeatures, and Myllostomella n.gen. described.2008040213有 孔 虫 Cornuspiroides 的 演 化 和 两 极 分 布 -利 用 形 态 学 特 征 和 简 约 性 理 论 来 推 断 系 统 发生 = Evolutin and bipolar distribution of Cornuspiroides(foraminifera): phylogenetic inferenceusing morphological characters and parsimony.( 英 文 ). Gudmundsson G. Journal of ForaminiferalResearch, 2002, 32(3): 308-318 3 图版 .The evolutionary relationship of three speciesbelonging to the genus Cornuspiroides is inferredusing four species of Cornuspira and themonotypic genus Cornuspirella as an outgroup.All known morphological features of the test arecoded as independent characters (synapomorphies)and their evolutionary relationship is inferredusing parsimony. The genus Cornuspiroidesappears as a monophyletic group on theresulting evolutionary trees, although evidentialsupport is weak (54%). The species Cornuspiroidesstriolatus and Cornuspiroides rotundusare endemics of the shallow and cold waters ofthe Arctic and the Antarctic seas, respectively.These two endemic species are found to share animmediate common ancestry, although supportis low (61%). The closest relative of the two polarspecies is inferred to be the little known Cornuspiroidesprofundum, which has been foundliving in the cold and shallow waters of the Arcticand also the temperate waters of the NorthAtlantic. Limitations of available data precludeC. profundum from being designated as theiractual ancestor. The current classificatory groupsof Cornuspiroidinae, Cornuspira, and Cornuspirinaeappear as artificial paraphyletic groupson the inferred tree and have no significant evidentialsupport of characters.2008040214利 用 有 孔 虫 的 同 位 素 参 数 对 上 部 水 团 的 分 层研 究 = A foraminiferal isotopic proxy for upperwater mass stratification. ( 英 文 ). Pak D K; KennettJ P. Journal of Foraminiferal Research,2002, 32(3): 319-327Modern oceanographic investigations showthat surface ocean warming is associated withincreased thickness of the mixed layer, deepeningof the thermocline, and reduction of upwellingstrength. These changes can profoundly affectsurface ocean biological productivity. Proxiesto measure past changes in upper water massstructure and stability are often poorly constrained.Stable oxygen isotope studies of planktonicforaminifera collected in sediment traps inSanta Barbara Basin, Southern California, demonstratethat the 18O difference (18O) betweenG. bulloides and N. pachyderma (d.) closelymonitors changes in the depth of the thermoclineand related thickness and stability of the mixedlayer. This proxy can be employed in investigationsof past changes in sea surface temperature,upwelling and thermocline strength on the Californiamargin, an area of great sensitivity toglobal change.2008040215根 据 ssu rRNA 基 因 序 列 推 断 出 的Allogromiid 有 孔 虫 的 系 统 发 生 = Phylogenyof Allogromiid foraminifera inferred from ssurRNA gene sequences. ( 英 文 ). Pawlowski J;Holzmann M; Berney C; Fahrni J; Cedhagen T;Bowser S S. Journal of Foraminiferal Research,2002, 32(4): 334-34368
Allogromiids are classically defined as agroup of monothalamous, soft-walled foraminiferans.Recent morphological, cytological, andmolecular studies, however, challenge this view,showing that the soft-walled allogromiids areclosely related to naked athalamids and unilocularagglutinated foraminiferans. To establish thephylogenetic relationships among these threegroups we obtained partial small-subunit ribosomalDNA sequences of 50 species and undeterminedmorphotypes, and compared them toother foraminiferal taxa. Phylogenetic analysesof our data show that allogromiids, athalamidsand astrorhizids comprise an assemblage of 13lineages branching together at the base of theforaminiferal tree. Among these lineages, twoare represented by a single species and fourcomprise similar genera, while the remainingseven are heterogeneous groups composed ofseveral species having different types of wallstructure and different test morphologies. Alllineages are relatively well supported, yet therelationships among them are not resolved. Inview of our data, we propose to revise the definitionof allogromiids to include all naked and testateunilocular granuloreticuloseans that divergedearly in the evolution of Foraminifera.2008040216日 内 瓦 湖 淡 水 有 孔 虫 类 的 过 去 和 现 在 =Freshwater foraminiferans from Lake Geneva:past and present. ( 英 文 ). Holzmann M; PawlowskiJ. Journal of Foraminiferal Research,2002, 32(4): 344-350 1 图 版 .Allogromiid foraminiferans are distributedover a wide range of marine habitats, but someare also reported from freshwater environments.Most of these descriptions date back to the 19thcentury. Among them, five species of freshwaterforaminiferans have been described and redescribedfrom Lake Geneva by the Swiss protozoologistEugène Penard. The different foraminiferalspecies were classified by Penard inthe genus Gromia, which at that time includedboth, filosean and granuloreticulosan species andwere later grouped by De Saedeleer in two allogromiidgenera, Allelogromia and Diplogromia.We collected several sediment samplesfrom Lake Geneva and from two other bodiesof freshwater in Switzerland and investigatedthem subsequently under a stereomicroscope.None of the forms described by Penard wereobserved. On the other hand, we have obtainedforaminiferal DNA sequences from DNA extractsof the same sediment samples. Phylogeneticanalysis of these sequences shows that theybranch with a clade of saccamminid foraminiferans,represented by the genera Ovammina andCribrothalammina. This is in agreement with theobserved morphological similarity betweenPenard’s species and the marine saccamminidsused in our study. The possibility that our sequencesbelong to some undetermined nakedforaminiferans, however, cannot be excludedand the molecular identification of the speciesdescribed by Penard must await the confirmationby sequencing DNA of isolated specimens.2008040217瑞 典 西 海 岸 一 个 大 的 单 房 室 有 孔 虫 新 属 新 种Toxisarcon synsuicidica = Toxisarcon synsuicidican. gen., n. sp., a large monothalamousforaminiferan from the west coast of Sweden.( 英 文 ). Cedhagen T; Pawlowski J. Journal ofForaminiferal Research, 2002, 32(4): 351-357 3图 版 .A new foraminiferan, Toxisarcon synsuicidican. gen., n. sp., is reported from clay bottoms ofKosterfjorden (Sweden). The species is characterizedby a large irregular cell body covered byan organic lining to which foreign particles areattached. It resembles an irregular clump of detritusor sediment aggregation and it can thereforeeasily be overlooked. Marine biologists whohave worked intensively with benthic fauna inthe area have never observed it despite the factthat it is quite common. The peculiar feature ofthe new species is its capacity to stock toxicproducts that can be harmful to itself if accidentallyreleased.2008040218英 国 苏 格 兰 西 海 岸 Linnhe 海 湾 新 种Toxisarcon alba 的 系 统 学 和 生 态 学 = Taxonomyand ecology of Toxisarcon alba, sp. nov.from loch Linnhe, west coast of Scotland, UK.( 英 文 ). Wilding T A. Journal of ForaminiferalResearch, 2002, 32(4): 358-363 2 图 版 .Toxisarcon alba n. sp., is a giant, naked marineforaminiferan recorded at two locations onthe west coast of Scotland, UK. Individuals werefound buried or semi-buried in well-oxygenatedsilty sand at 10 to 30m water depth. A distinctive,slightly raised mound of 18–40mm diameter,through which there were many perforations,indicated the presence of an individual on theseabed. Partial removal of the overlying sedimentrevealed the naked opaque, white, stickycell body. Toxisarcon alba exists in three distinctforms. In the most commonly occurringform (and the only form observed in the field),the central portion of the cytoplasm extends peripherallyinto numerous large cytoplasmic extensionsand reticulopodia (branched form). Inaquaria, some individuals transformed into aconsolidated form (spheroid form) by retractingmost or all of the reticulopods resulting in a cellwith a very well defined edge. On four occasionsa third very diffuse form, resembling a fungal69
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第 4 卷 第 4 期古 生 物 学
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Fossils, i.e. remains of living org
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nus, a new ichnospecies of Fuersich
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以 松 柏 类 和 盾 籽 类 为
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Island, Papua New Guinea. ( 英 文
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A C. Canadian Journal of Earth Scie
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mated depositional interval of 10 t
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北 海 古 近 纪 一 种 新 型
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velopment of both the sedimentary a
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China. ( 英 文 ). Huang Diying; N
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德 国 南 部 上 侏 罗 统 (Ox
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shelf basin off the northern coast
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like base referred to Primocandelab
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鱼 类2008040413大 西 洋 鳕
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long as the humerus plus radius, ve
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members in both size and abundance
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ple's Republic of China. ( 英 文
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K-P transition in Brazil. ( 英 文
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wide tooth crowns, a sub-rectangula
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2008040459罗 马 尼 亚 特 兰
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logical descriptions of all availab
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东 副 特 提 斯 晚 萨 马 期
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S; Vaiani S C. Rivista Italiana di
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adiocarbon dating the arrival of th
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able problems, particularly differe
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done with glyphs on paper. In this
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obtained through extensive field-ba
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noli G; Maletz J; Stouge S. Canadia
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法 国 蒙 塔 格 尼 努 尔 中
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presence of well-rounded, coarse-sa
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缅 甸 北 部 Hukawng 峡 谷 琥
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graphy and Geological Correlation,
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plex,northwestern Iran. ( 英 文 )
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松 辽 盆 地 晚 白 垩 世 青
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ous nannoplankton) points of view.O
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the"Yanshan Movement".During the Mi
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geochemical approach discussed in t
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sediments from the northern shelf o
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意 大 利 北 亚 平 宁 Lamone
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nano record further into the early
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lankovitch-scale frequency. Analyse