A three-year sediment trap experiment tostudy the production and flux of planktonic foraminiferawas conducted in the Cariaco Basin,Venezuela, between January, 1997, and December,1999. The results allow direct comparisonbetween planktonic foraminiferal flux and assemblagecomposition, and the climatology andhydrography of the region on seasonal and interannualtime scales. The Cariaco Basin is locatedon the northern continental shelf of Venezuela ina region characterized by intense upwelling dueto the seasonal migration of the Inter-TropicalConvergence Zone (ITCZ) and associated variationsin tradewind intensity. Hydrographic observationsindicate that coastal upwelling occursduring the late winter-spring at which time seasurface temperatures are lowest (~22°C) andprimary productivity is high. Immediately followingmaximum primary production the totalflux of planktonic foraminifera increases fromless than 100 shells m–2 day–1 to 4000–8000shells m–2 day–1. Approximately half of thetotal annual flux is produced during this period(February–May). Nine species, or varieties, ofplanktonic foraminifera constitute >85% of theassemblages: Orbulina universa, Globigerinoidesruber (pink and white varieties), Globigerinabulloides, Globigerina quinqueloba,Neogloboquadrina dutertrei, Globorotalia crassaformis,Globorotalia menardii, and Globigerinitaglutinata. These species are present yearround;however, their flux and contribution tothe population vary seasonally and interannually.The peak annual flux of all nine species occursduring spring upwelling. However, Globigerinabulloides is dominant during this period, withvalues reaching ~4000 shells m–2 d–1, or 50 to75% of the total flux during upwelling. Sea surfacetemperatures were estimated for both sedimenttrap and core-top assemblages using theModern Analog Technique (MAT) and comparedwith conductivity-temperature-depth(CTD) data to evaluate the accuracy of thismethod for reconstructing paleotemperatures inCariaco Basin. The assemblage temperature estimatesare within ~1–3°C of the measured temperatures.2008040183意 大 利 撒 丁 岛 Bathonian 地 区 Hormosinidae科 Lituolida 亚 科 的 一 新 属 新 种 Posadia feroniensis= Posadia feroniensis n. gen.,n.sp.(Lituolida,Hormosinidae)from the Bathonianof Sardinia, Italy. ( 英 文 ). Giusberti L; CoccioniR. Journal of Foraminiferal Research, 2003,33(3): 211-218 4 图 版 .Detailed stratigraphical and paleobiologicalstudies on the Mesozoic carbonate platform ofeastern Sardinia (Italy) revealed the occurrenceof an unknown foraminifer in the Bathonian(Middle Jurassic) Dorgali Dolostone croppingout in the Posada hill. Posadia feroniensis n.gen., n. sp., a new distinctive agglutinated foraminiferreferable to the family HormosinidaeHaeckel, 1894, is formally described and figured.Posadia is morphologically characterized by anuniserial, roughly finished test with inverted"V"-like chambers in longitudinal section.2008040184巴 勒 斯 坦 印 度 盆 地 苏 来 曼 山 脉 始 新 世 浮 游 有孔 虫 的 生 物 地 层 学 = Eocene planktic foraminiferalbiostratigraphy of the Sulaimanrange,Indus basin,Pakistan. ( 英 文 ). Warraich MY; Nishi H. Journal of Foraminiferal Research,2003, 33(3): 219-236 2 图 版 .Paleocene and Eocene marine sequences distributedin Rakhi Nala and Zinda Pir Anticlineareas of the eastern part of the Sulaiman Rangein Pakistan record the geological history andtectonic events during the last stage of collisionbetween the Indian and Eurasian Plates and closureof the Tethyan Ocean. The Eocene sequencesconsist of siltstone and mudstone withsome limestone intercalations, and are dividedinto the Shaheed Ghat, Baska and Kirthar Formationsin ascending order. We studied the Eocenebiostratigraphy of planktic foraminifers inthis region to reveal the geological ages of theEocene strata based on (international) correlationwith standard tropical/subtropical zones. Theplanktic foraminiferal assemblages are composedof 78 species of 19 different genera, includingmany zonal markers of tropical to subtropicalregions. We identified 8 biostratigraphiczones, Zone P7, Acarinina pentacamerata PartialRange Zone, Planorotalites palmerae TotalRange Zone, A. soldadoensis Assemblage Zone,Zone P12, Zone P13, Orbulinoides beckmanni-Truncorotaloides rohri Interval Zone, and Truncorotaloidesrohri-Turborotalia cerroazulensiscunialensis Interval Zone in ascending order.These zones correspond to Zones P7 to P15 oftropical/subtropical schemes. However, the upperpart of Zone P10 to Zone P11 is not recognizedbecause of an interval that is barren ofplanktic foraminifers in the Baska Formation.The geological age of the Shaheed Ghat Formationis assigned to the Early Eocene throughearly Middle Eocene (Zones P7 to P10,52.3~45.8 Ma). The Baska Formation is apparentlyequivalent to Zone P10 (early Middle Eocene),although its direct age is uncertain as it isbarren of planktic foraminifers. The fossiliferousmicritic limestone forming the lowermost part ofthe Kirthar Formation yielded recrystallizedplanktic foraminifers and calcareous nannoplanktonindicating an early Middle Eocene agespanning calcareous nannoplankton Zones NP14to NP15 (48.5~43.4 Ma), while the upper part of58
the Kirthar Formation ranges in age from middleMiddle Eocene to early Late Eocene (Zones P12to P15, 43.6~35.2 Ma). Hence the strata of theKirthar Formation spans in age from early MiddleEocene through early Late Eocene based onrecognition of the planktic foraminiferal ZonesP10 to P15 (49~35.2 Ma).2008040185始 新 世 浮 游 有 孔 虫 Hantkenina 的 起 源 和 形 态学 = Origin and morphology of the Eoceneplanktonic foraminifer Hantkenina. ( 英 文 ). CoxallH K; Huber B T; Pearson P N. Journal ofForaminiferal Research, 2003, 33(3): 237-261 9图 版 .Study of the origin and early evolution of thetubulospine-bearing planktonic foraminiferalgenus Hantkenina reveals that it evolved graduallyfrom the clavate species Clavigerinella eocanicain the earliest middle Eocene and is unrelatedto the genus Pseudohastigerina.Clavigerinella eocanica and the lower middleEocene species Hantkenina nuttalli share manymorphologic features and show similar developmentalpatterns but differ significantly inthese aspects from P. micra. Rare, transitionalClavigerinella-Hantkenina forms from the Helvetikumsection of Austria bridge the gap betweenclavate and tubulospinose morphologies,providing direct, stratigraphically-ordered evidenceof the evolutionary transition betweenHantkenina and Clavigerinella. Clavigerinellidancestry is traced to a previously undescribedlow-trochospiral species, Parasubbotina eoclavasp. nov., at Ocean Drilling Program Site 865.We speculate that Hantkenina originated throughcompetition for limited food resources in a deep,oxygen-deficient habitat below the thermocline.The tubulospines may represent a structural adaptationto this new trophic strategy, allowingthe organism to harvest a greater volume of waterat minimal metabolic cost. The abrupt occurrenceof Hantkenina in pelagic sediment coresfrom the central Pacific and other regions of theworld ocean may represent immigration intothese areas following speciation within the hydrographicallyevolving Tethys Seaway. Alternatively,cladogenesis may have occurred over awider area, but due to a contemporaneous globalhiatus the fossil record of this bioevent is poorlypreserved.2008040186分 子 生 物 学 数 据 揭 示 的 有 孔 虫 Nummulitid科 的 平 行 演 化 = Molecular data reveal parallelevolution in Nummulitid foraminifera. ( 英 文 ).Holzmann M; Hohenegger J; Pawlowski J.Journal of Foraminiferal Research, 2003, 33(4):277-284 1 图 版 .Nummulitidae are the largest extant calcareousForaminifera, and are widely distributed intropical and subtropical shallow-water seas.Classical morphology-based taxonomy dividesthe Nummulitidae in two subfamilies, the Nummulitinaeand Heterostegininae, according to thepresence or absence of secondary septa. To testthe evolutionary importance of this morphologicalfeature, phylogenetic relationships of fiveRecent nummulitid genera were investigated bysequencing fragments of the SSU and LSUrRNA gene. According to our results, speciescharacterized by septate chambers (Heterosteginadepressa, Planostegina operculinoides, andCycloclypeus carpenteri) either group with specieslacking septate chambers (Operculina ammonoides,Nummulites venosus) or branch separately.This suggests that chamber subdivisionsdeveloped several times independently in theevolutionary history of the Nummulitidae, providingan example of parallel evolution in Foraminifera.2008040187得 克 萨 斯 科 罗 拉 多 河 生 境 破 坏 的 有 孔 虫 响 应= Foraminiferal response to habitat disruption:Arroyo Colorado, Texas. ( 英 文 ). Buzas-Stephens P; Pessagno E A. Journal of ForaminiferalResearch, 2003, 33(4): 294-308 1 图版 .In order to assess the possible effects of pollutantson foraminifera along the Texas GulfCoast, sediment cores were taken in four areasconsidered to be at risk: southern Laguna Madre,Nueces Bay, the Arroyo Colorado, and LagunaAtascosa. Geochemical analyses indicate that thesediments are relatively uncontaminated. Foraminiferalanalyses, however, reveal otherwiseundetected inconsistencies and changes withtime. First, though foraminiferal assemblages inthe surface sediments of Laguna Madre, NuecesBay, and Laguna Atascosa are typical, they areatypical in the Arroyo Colorado. In Texas estuarineenvironments, Ammonia often comprises alarge proportion of the assemblage. But in theArroyo Colorado, an estuarine river, the assemblageis almost all Elphidium to the exclusion ofAmmonia. A second observation is that downcorein the Arroyo Colorado cores, Ammoniabecomes abundant as numbers of El-phidiumdecrease. The assemblage at the base of the ArroyoColorado cores is thus more typical for theGulf Coast, while that in the surface sediments isnot. This down-core change in the foraminiferalassemblage is coincident with a change in sedimentgrain size, both most likely due to thedredging of the Harlingen ship channel in thelate 1940s. The reasons why Ammonia, an opportunisticgenus that thrives in a wide range ofenvironmental conditions, was unable to tolerate59
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第 4 卷 第 4 期古 生 物 学
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Fossils, i.e. remains of living org
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nus, a new ichnospecies of Fuersich
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Brachyelasma, Amplexobrachyelasma,
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litic biozones lower than their FAD
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The middle Permian Cryptospirifer f
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石 科 = Ammonoid faunas from the
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Buchiid bivalves are geographically
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velopment of both the sedimentary a
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China. ( 英 文 ). Huang Diying; N
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德 国 南 部 上 侏 罗 统 (Ox
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shelf basin off the northern coast
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like base referred to Primocandelab
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鱼 类2008040413大 西 洋 鳕
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long as the humerus plus radius, ve
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members in both size and abundance
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ple's Republic of China. ( 英 文
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K-P transition in Brazil. ( 英 文
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wide tooth crowns, a sub-rectangula
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2008040459罗 马 尼 亚 特 兰
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logical descriptions of all availab
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东 副 特 提 斯 晚 萨 马 期
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S; Vaiani S C. Rivista Italiana di
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adiocarbon dating the arrival of th
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able problems, particularly differe
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done with glyphs on paper. In this
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obtained through extensive field-ba
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noli G; Maletz J; Stouge S. Canadia
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presence of well-rounded, coarse-sa
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缅 甸 北 部 Hukawng 峡 谷 琥
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graphy and Geological Correlation,
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plex,northwestern Iran. ( 英 文 )
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ous nannoplankton) points of view.O
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sediments from the northern shelf o
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nano record further into the early
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lankovitch-scale frequency. Analyse