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Assessment, Conservation and Sustainable Use of Forest Biodiversity

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<strong>Assessment</strong>, <strong>Conservation</strong> <strong>and</strong> <strong>Sustainable</strong> <strong>Use</strong> <strong>of</strong> <strong>Forest</strong> <strong>Biodiversity</strong><br />

28<br />

SUSTAINABLE FOREST MANAGEMENT AUGMENTS DIVERSITY OF<br />

VASCULAR PLANTS IN GERMAN FORESTS<br />

G.von Oheimb*, H. Ellenberg**, J. Heuveldop***, W.U. Kriebitzsch**<br />

*Institute for Ecology <strong>and</strong> Environmental Chemistry, University <strong>of</strong> Lueneburg, Germany;<br />

** Institute for World <strong>Forest</strong>ry, Federal Research Centre for <strong>Forest</strong>ry <strong>and</strong> <strong>Forest</strong> Products, Hamburg, Germany;<br />

*** Chair for World <strong>Forest</strong>ry, University <strong>of</strong> Hamburg, Germany. –<br />

E-mail: ellenberg@holz.uni-hamburg.de<br />

Keywords: species diversity, vascular plants, forest management intensity, sustainable forest management Germany<br />

Introduction<br />

There exists a need in politics, administration, <strong>and</strong> in practice <strong>of</strong> forest management for knowledge <strong>and</strong><br />

decision support information on conservation <strong>and</strong> sustainable use <strong>of</strong> biological diversity in German forests.<br />

Therefore the present study investigated the following hypotheses:<br />

1) As a consequence <strong>of</strong> forest management, there exists a higher number <strong>of</strong> vascular plant species per unit<br />

area in the herb layer <strong>of</strong> managed forests in relation to ecologically comparable non managed forests.<br />

2) With growing management intensity, species composition in vascular plants shifts towards higher<br />

fractions <strong>of</strong> indicator species for disturbance.<br />

Methods<br />

Vegetation analysis was conducted in 1997 <strong>and</strong> 1998 in broadleaved lowl<strong>and</strong> forests <strong>of</strong> Northern Germany on<br />

rich soils (county <strong>of</strong> Herzogtum Lauenburg, east <strong>of</strong> Hamburg: “SH”) as well as on moderately rich soils<br />

(beech, Fagus silvatica, forests <strong>of</strong> the Mueritz National Park “MV”). Virgin forests do not exist in Germany. So,<br />

non-managed (at least for 30 years, up to more than 150 years) forests were contrasted against managed forests<br />

<strong>of</strong> different management intensity. The latter was measured among others as cubic metres <strong>of</strong> timber taken per<br />

hectare <strong>and</strong> per year. Soil quality <strong>and</strong> water supply as well as tree species composition <strong>and</strong> st<strong>and</strong> age were as<br />

comparable as possible. Contrary to no longer managed forests, thinning <strong>and</strong> final timber harvest were<br />

conducted in managed forests.<br />

Results<br />

In almost every comparison, higher mean species numbers per unit area were found in managed forests as<br />

opposed to no longer managed forests (Fig. 1 <strong>and</strong> 2).<br />

In every investigated forest st<strong>and</strong>, managed or not, the widespread <strong>and</strong> frequent indicator species for the forest<br />

types <strong>of</strong> Querco-Fagetea <strong>and</strong> Fagetalia (“typical” forest species in a narrow sense)existed in similar densities. On<br />

the other h<strong>and</strong>, a significantly higher distribution <strong>and</strong> density was observed in managed forests for nitrophytes<br />

<strong>and</strong> for indicator species for light (open canopy) <strong>and</strong> for soil compaction (Tab. 1 <strong>and</strong> 2). The steadyness values<br />

<strong>of</strong> these latter species grows further with raising intensity <strong>of</strong> forest management (Tab. 1, Fig. 1).<br />

Conclusions<br />

Higher numbers <strong>of</strong> vascular plant species per unit area are possible in managed forests due to a better light<br />

climate in the herb layer as a consequence <strong>of</strong> forest operations for timber harvest that open up the canopy.<br />

Additionally, heterogeneous soil conditions (bare, accumulated, compacted, etc.) are caused by timber moving<br />

operations <strong>and</strong> a higher forest road density, both leading to better conditions for establishment <strong>and</strong> growth <strong>of</strong><br />

vascular plants at least temporarily.<br />

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