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Introduction to the Modeling and Analysis of Complex Systems

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54 CHAPTER 4. DISCRETE-TIME MODELS I: MODELINGf(x)af(x) = –a – 1Kx + a10KxFigure 4.5: The simplest example <strong>of</strong> how <strong>the</strong> growth ratio a = f(x) should behave asa function <strong>of</strong> population size x.this case, <strong>the</strong> equation becomes x t = 0, so <strong>the</strong>re is no growth. This makes perfect sense;if <strong>the</strong>re are no organisms left, <strong>the</strong>re should be no growth. Ano<strong>the</strong>r extreme case: Whathappens when x t−1 = K? In this case, <strong>the</strong> equation becomes x t = x t−1 , i.e., <strong>the</strong> systemmaintains <strong>the</strong> same population size. This is <strong>the</strong> new convergent behavior you wanted <strong>to</strong>implement, so this is also good news. Now you can check <strong>the</strong> behaviors <strong>of</strong> <strong>the</strong> new modelby computer simulations.Exercise 4.9 Simulate <strong>the</strong> behavior <strong>of</strong> <strong>the</strong> new population growth model for severaldifferent values <strong>of</strong> parameter a <strong>and</strong> initial condition x 0 <strong>to</strong> see what kind <strong>of</strong>behaviors are possible.(x t = − a − 1 )Kx t−1 + a x t−1 (4.23)For your information, <strong>the</strong> new model equation we have just derived above actuallyhas a particular name; it is called <strong>the</strong> logistic growth model in ma<strong>the</strong>matical biology <strong>and</strong>several o<strong>the</strong>r disciplines. You can apply a parameter substitution r = a − 1 <strong>to</strong> make <strong>the</strong>

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