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Introduction to the Modeling and Analysis of Complex Systems

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4.6. BUILDING YOUR OWN MODEL EQUATIONS WITH MULTIPLE VARIABLES 57PreyPopulation : xNaturally grows<strong>to</strong> carrying capacityif isolatedPositive influencePreda<strong>to</strong>rs’ growth rateis higher withmore prey+-Negative influencePrey’s survival rate+ -is lower withmore preda<strong>to</strong>rsPreda<strong>to</strong>rsPopulation : yNaturally decaysif isolatedFigure 4.7: Interactions between <strong>the</strong> prey <strong>and</strong> preda<strong>to</strong>r populations illustrated in acausal loop diagram.The inherent dynamics <strong>of</strong> <strong>the</strong> two variables are quite straightforward <strong>to</strong> model. Sincewe already know how <strong>to</strong> model growth <strong>and</strong> decay, we can just borrow those existingmodels as building components, like this:x t = x t−1 + r x x t−1 (1 − x t−1 /K) (4.28)y t = y t−1 − d y y t−1 (4.29)Here, I used <strong>the</strong> logistic growth model for <strong>the</strong> prey (x) while using <strong>the</strong> exponential decaymodel for <strong>the</strong> preda<strong>to</strong>rs (y). r x is <strong>the</strong> growth rate <strong>of</strong> <strong>the</strong> prey, <strong>and</strong> d y is <strong>the</strong> death rate <strong>of</strong><strong>the</strong> preda<strong>to</strong>rs (0 < d y < 1).To implement additional assumptions about <strong>the</strong> preda<strong>to</strong>r-prey interactions, we need <strong>to</strong>figure out which part <strong>of</strong> <strong>the</strong> equations should be modified. In this example it is obvious,because we already know that <strong>the</strong> interactions should change <strong>the</strong> death rate <strong>of</strong> <strong>the</strong> prey<strong>and</strong> <strong>the</strong> growth rate <strong>of</strong> <strong>the</strong> preda<strong>to</strong>rs. These terms are not yet present in <strong>the</strong> equationsabove, so we can simply add a new unknown term <strong>to</strong> each equation:x t = x t−1 + rx t−1 (1 − x t−1 /K) − d x (y t−1 )x t−1 (4.30)y t = y t−1 − dy t−1 + r y (x t−1 )y t−1 (4.31)Now <strong>the</strong> problems are much better defined. We just need <strong>to</strong> come up with a ma<strong>the</strong>maticalform for d x <strong>and</strong> r y .

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