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Soybean and Bees

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Trichomes <strong>and</strong> nectaries<br />

Gynoecium non-secretory <strong>and</strong> secretory trichomes are associated with nectary development.<br />

During the final period of preactive nectary development, three types of trichomes<br />

appears on the epidermis of the gynoecium. The first type is elongated, thinwalled,<br />

non- gl<strong>and</strong>ular, unicellular trichome, containing a large vacuole with peripheral<br />

cytoplasm <strong>and</strong> a very large nucleus (Horner et al., 2003).<br />

The second type of elongated, non-gl<strong>and</strong>ular trichome consists of one or two basal cells<br />

with thickened walls <strong>and</strong> a long terminal cell with a much thicker wall than the unicellular<br />

trichome. The outer trichome wall also displays papillae. These trichomes contain<br />

relatively large nuclei with large nucleoli <strong>and</strong> occur in the uppermost part of the gynoecium<br />

toward the stigma (Horner et al., 2003).<br />

The third type of trichome develops later than the two elongated, non-gl<strong>and</strong>ular trichomes,<br />

is smaller <strong>and</strong> shorter, consisting of 5-7 cells in linear array, as described by<br />

Horner et al. (2003). These latter trichomes are dispersed among the longer trichomes<br />

from the base to the upper part of the gynoecium. The nucleus of each cell is much<br />

smaller than the nucleus in the two types of non-gl<strong>and</strong>ular, elongated trichomes, <strong>and</strong> it<br />

is centrally located in each cell.<br />

SoybeAn <strong>and</strong> bees<br />

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