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Wildlife of Lao PDR: 1999 Status Report - IUCN

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<strong>Wildlife</strong> <strong>of</strong> <strong>Lao</strong> <strong>PDR</strong>: <strong>1999</strong> <strong>Status</strong> <strong>Report</strong><br />

[• Nycticebus pygmaeus Pygmy Loris (= Lesser Slow<br />

Loris M5 ; = Pygmy Slow Loris M4 )]. Conservation Significance:<br />

Globally Threatened - Vulnerable; Little Known in <strong>Lao</strong> <strong>PDR</strong>;<br />

CITES Appendix II. Endemic to parts <strong>of</strong> <strong>Lao</strong> <strong>PDR</strong>, southern<br />

Yunnan (China), Vietnam and Cambodia; occurs only east <strong>of</strong><br />

the Mekong (Corbet and Hill 1992). Documented Range and<br />

Habitat: North M13 , centre M8 , south M14 ; all provisional. Evergreen<br />

forest in the Annamites and other hilly areas. <strong>Status</strong><br />

Information: Field records <strong>of</strong> small lorises from a variety <strong>of</strong><br />

areas (Table 12); not yet found commonly anywhere above<br />

500 m. Although described as ‘common’ throughout <strong>Lao</strong> <strong>PDR</strong><br />

(Delacour 1940), there are few specific historical records:<br />

Phongsali, Ban Thateng on the Bolaven Plateau (Osgood<br />

1932) and Savannakhet (Bourret 1944). Taxonomic issues:<br />

Recent research in Bolikhamxai Province indicates that at<br />

least three species <strong>of</strong> loris inhabit <strong>Lao</strong> <strong>PDR</strong> (Alterman and<br />

Freed 1997). To which, if any, the name pygmaeus is applicable<br />

is unclear (L. Alterman in litt. 1998). The publication<br />

<strong>of</strong> sightings as N. pygmaeus by Duckworth (1994b) was premature,<br />

in omitting any comparative examination with the<br />

type specimen <strong>of</strong> pygmaeus. Nonetheless, the general conclusion<br />

holds, that two lorises <strong>of</strong> different sizes are ecologically<br />

sympatric in Phou Xang He NBCA. Similar patterns<br />

have been found widely elsewhere, from Bolaven Northeast<br />

PNBCA in the south, to Nam Kading NBCA in the north<br />

(Evans et al. in prep. b). Until the issue is clarified, it is best<br />

to consider the named species N. pygmaeus as Little Known<br />

in <strong>Lao</strong> <strong>PDR</strong>, while accepting that it may turn out to be<br />

common and widespread and <strong>of</strong> no immediate conservation<br />

concern.<br />

[ Nycticebus intermedius Intermediate Loris (= Intermediate<br />

Slow Loris M5 )]. Context: May occur in north-east <strong>Lao</strong> <strong>PDR</strong>,<br />

but apparently known only from the type from Vietnam and<br />

various specimens from Yunnan, China (Corbet and Hill<br />

1992). Special Significance: CITES Appendix II. Speculated<br />

to be endemic to a small area around the <strong>Lao</strong>-Vietnam-Yunnan<br />

(China) border (Corbet and Hill 1992). Taxonomic issues:<br />

The type was collected in northern Vietnam (Dao Van Tien<br />

1960). N. intermedius has been considered synonymous with<br />

N. pygmaeus on grounds <strong>of</strong> both morphology and genetics<br />

(Groves 1971, Zhang et al. 1993). There are more than two<br />

species <strong>of</strong> loris in <strong>Lao</strong> <strong>PDR</strong> (Alterman and Freed 1997; L.<br />

Alterman in litt. 1998), but whether intermedius is a valid<br />

name, and whether any <strong>of</strong> the forms in <strong>Lao</strong> <strong>PDR</strong> belong to it,<br />

are not yet clear.<br />

<strong>Status</strong> Information on Lorises: Lorises are not favoured as<br />

food in <strong>Lao</strong> <strong>PDR</strong> (e.g. Duckworth 1994b) but some people<br />

are not averse to eating them (e.g. Davidson 1998). They are<br />

kept as pets (Salter 1993a; Plate 13) and used in medicinal<br />

preparations (Baird 1995b). The latter may be the most significant<br />

threat. Such harvesting goes on at high levels in neighbouring<br />

Cambodia (Timmins and Men Soriyun 1998) and<br />

176<br />

Vietnam (Ratajszczak 1988). KK has received reports <strong>of</strong> the<br />

export to Vietnam <strong>of</strong> large numbers from Bolikhamxai and<br />

Khammouan Provinces. Animals are caught in various ways;<br />

the most damaging is when trees are cut down so that the<br />

loris can be captured, which occurs at least in Bolikhamxai<br />

Province (I. Johnson verbally <strong>1999</strong>).<br />

Conservation Management and Research Proposed for<br />

Lorises:<br />

• Clarification <strong>of</strong> the number <strong>of</strong> forms present and their<br />

geographical and ecological distributions, to allow identification<br />

<strong>of</strong> taxa in need <strong>of</strong> specific action, and the actions<br />

themselves.<br />

• Pending this, collection <strong>of</strong> detailed notes on appearance<br />

<strong>of</strong> animals (supported whenever possible with photographs<br />

and measurements) seen on field surveys or in<br />

markets or villages to allow possible retrospective identification.<br />

• Investigation <strong>of</strong> trade threat, with due attention to international<br />

issues.<br />

• Enactment and enforcement <strong>of</strong> a nocturnal hunting ban<br />

in NBCAs within the range <strong>of</strong> forms identified as threatened<br />

in <strong>Lao</strong> <strong>PDR</strong>; immediate enactment <strong>of</strong> such a ban in<br />

core zones <strong>of</strong> all NBCAs.<br />

Cercopithecidae: Old-world monkeys (9 species in <strong>Lao</strong><br />

<strong>PDR</strong>; 81 worldwide)<br />

• Macaca nemestrina Pig-tailed Macaque (= Pigtail<br />

Macaque M4 ). Conservation Significance: Globally Threatened<br />

- Vulnerable; Potentially At Risk in <strong>Lao</strong> <strong>PDR</strong>; CITES Appendix<br />

II. Documented Range and Habitat: North M9 , centre M8 ,<br />

south M8 . Evergreen forest, and also in mosaic with more open<br />

habitats such as pine and dry dipterocarp forest; lowlands<br />

and lower hills, and locally in mountains. <strong>Status</strong> Information:<br />

See below. Recorded in many survey areas (Table 12),<br />

in several <strong>of</strong> which it was relatively common, including some<br />

highly degraded regions. Estimated group sizes in <strong>Lao</strong> <strong>PDR</strong><br />

rarely exceeded 20-30. In some areas <strong>of</strong> other countries (e.g.<br />

Khao Yai National Park, Thailand; rural Dak Lak Province,<br />

Vietnam), probably those relatively undisturbed by opportunistic<br />

shooting, Pig-tailed Macaque groups number 50-150<br />

animals (Le Xuan Canh et al. 1997; A. R. Nettelbeck verbally<br />

1996). Only one recent record in <strong>Lao</strong> <strong>PDR</strong> specifically<br />

noted a group <strong>of</strong> such size (over 100 in Dong Phou Vieng<br />

NBCA; Steinmetz 1998a).<br />

• Macaca assamensis Assamese Macaque (= Assam<br />

Macaque M4 ). Conservation Significance: Globally Threatened<br />

- Vulnerable; Potentially At Risk in <strong>Lao</strong> <strong>PDR</strong>; CITES Appendix<br />

II. Documented Range and Habitat: North M15 ,<br />

centre M13 , south M8 . Evergreen forests. Usually the commonest<br />

macaque in hills and mountains, occurring down to 300

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