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XIV iNinoDtrcTiox.<br />
chiefly owing to the Lack of properly preserved material fgtr<br />
iuvestigatiou ; it may not, however, be out of place to here insert<br />
a few binomic notes which concern the families generally, these<br />
being grouped under the various headings as below :—<br />
1. Breeding Habits.<br />
2. Larval Stages.<br />
3 Development from Post-Larval Stages.<br />
4. Habitat and Mode of Life.<br />
5. Movements, Locomotion, and Dispersal.<br />
6. Economic Uses.<br />
1. Breeding Habits.—Generally speaking the sexes in both the<br />
freshwater Gastropoda and Pelecypoda are distinct, though here<br />
and there examples of hermaphroditism are Jmown to exist, as,<br />
for instance, in the case of certain species of Anodonta.<br />
In the former case, the sexes can frequently be distinguished<br />
by the form of the shell, that of the female being of a larger size<br />
and generally more inflated than is that of the male. In the<br />
Gastropoda this is especially to be remarked in the Viviparida;,<br />
while in the Pelecypoda it is chiefly evident in the UnionidsE, the<br />
female in this group being noticeably broader than the male.<br />
In the fluviatile Gastropod families sexual contact usually takes<br />
place, ova as a result being de])Osited in the majority of instances,<br />
though in the case of the Tiaridse and Viviparidse, as the name of<br />
the latter would imply, the young are produced alive.<br />
In the Pelecypoda, however, union of the sexes does not take<br />
place, the male at certain seasons in the year freely discharging<br />
the spermatozoa into the water, these being introduced into the<br />
inhalent siphons of the female by means of currents set up by<br />
ciliary movements, fertilization taking place either in the oviduct<br />
itself or else in specialized spaces of the mantle cavity.<br />
The quantity of eggs or young produced by the different<br />
families varies enormously, this, while attaining in the Pelecypoda<br />
to thousands, and sometimes even to hundreds of thousands, as in<br />
the case of certain species of Unio and Anodonta, falls in Planorhis<br />
and Limncea to anything from twenty to a hundred, and in Ancylus<br />
to such a small total as five or six only, while Vivipara and Tiara<br />
average not more than about fifteen individuals at .a time.<br />
In Tiara the embryos are developed in a marsiipium which is<br />
formed by an infolding of the skin near the base of the right<br />
tentacle, while in the TJnionidse, CyrenidsB, and some other<br />
Pelecypoda development takes place in the spaces betweenlthe<br />
folds of the gills where, in the Cyrenidas, special marsupia exist<br />
for their reception'.<br />
2. Larval stages.—In the fluviatile Gastropoda at birtli the<br />
animal is generally more or less similar to that of the half-grown<br />
or adult state, though the shell differs considerably; in the earlier<br />
stages it is exogastric or coiled forward over the head of the<br />
animal, but rapidly assumes the normal spiral of the adult. In