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difficult to foresee and which reflect <strong>the</strong> <strong>of</strong>ten complex <strong>in</strong>ter-dependency <strong>of</strong> plants,<br />

animals and <strong>the</strong>ir physical environment (Hunter and Green, 1995).<br />

The negative impacts <strong>of</strong> tourists' feet are not restricted to terrestrial ecosystems.<br />

Hamele (1988) notes that <strong>the</strong> damage to coral reefs result<strong>in</strong>g from <strong>the</strong> contact <strong>of</strong><br />

divers' flippers and <strong>the</strong> chok<strong>in</strong>g effect <strong>of</strong> disturbed sediment can be severe. Blake<br />

and Becher (1997), referr<strong>in</strong>g to <strong>tourism</strong> <strong>in</strong> <strong>the</strong> Pacific Island States, report that a<br />

large proportion <strong>of</strong> <strong>the</strong> coral and small fish life around <strong>the</strong> marg<strong>in</strong>s <strong>of</strong> boat jetties and<br />

hotel beaches has been irreparably damaged or destroyed due to excessive walk<strong>in</strong>g<br />

on coral reef beds by tourists at low tide. The action <strong>of</strong> motor boats, surf boards and<br />

yachts also damages aquatic ecosystems, such as reed beds, while <strong>the</strong> construction<br />

<strong>of</strong> mar<strong>in</strong>as to house <strong>the</strong>se crafts can completely destroy, or degrade beyond repair,<br />

important mangrove and o<strong>the</strong>r lagoon habitats (Jackson, 1984).<br />

Tourism may also lead to <strong>the</strong> selective removal, collection or kill<strong>in</strong>g <strong>of</strong> (<strong>of</strong>ten rare)<br />

plants and animals. McLaren (1998), for example, argues that <strong>the</strong> <strong>natural</strong><br />

environment <strong>of</strong> some African countries is <strong>in</strong>creas<strong>in</strong>gly be<strong>in</strong>g destroyed through<br />

pressures from wildlife <strong>tourism</strong>. The collection and use <strong>of</strong> wild plants and animal<br />

species <strong>in</strong> countries like Senegal for <strong>the</strong> manufacture and market<strong>in</strong>g <strong>of</strong> trophies, gifts<br />

and souvenirs for <strong>the</strong> <strong>tourism</strong> <strong>in</strong>dustry has become a major problem. This is<br />

occurr<strong>in</strong>g even where <strong>the</strong> animals are 'protected', and contributes to <strong>the</strong><br />

development <strong>of</strong> ecological imbalances. The collection and kill<strong>in</strong>g <strong>of</strong> mar<strong>in</strong>e animals<br />

for <strong>the</strong> souvenir trade has also been highlighted, for example, <strong>in</strong> <strong>the</strong> Mediterranean<br />

and Pacific Islands (Milne, 1990), with marked impacts on local ecology. Hunter and<br />

Green (1995) describe <strong>the</strong> collection <strong>of</strong> shells, rocks and plants on <strong>the</strong> unique<br />

Galapagos Islands by poorly controlled groups <strong>of</strong> tourists from private yachts and<br />

small tourist boats <strong>of</strong>ten ventur<strong>in</strong>g beyond marked tourist routes. Also observed <strong>in</strong><br />

this study was <strong>the</strong> grow<strong>in</strong>g disturbance <strong>of</strong> plant and animal life close to tourist trails,<br />

lead<strong>in</strong>g to fears that animals would ultimately avoid certa<strong>in</strong> areas, as animals may<br />

well migrate out <strong>of</strong> an area used by tourists if harassed. Alternatively, <strong>the</strong> over­<br />

reliance <strong>of</strong> tourists on certa<strong>in</strong> animal species as a source <strong>of</strong> food may prove to be<br />

disadvantageous to <strong>the</strong> animals <strong>in</strong> <strong>the</strong> long term (S<strong>in</strong>diyo and Pertet, 1984).<br />

Dur<strong>in</strong>g certa<strong>in</strong> times <strong>of</strong> <strong>the</strong> year, tourists may have a severe impact on animal<br />

breed<strong>in</strong>g success, and <strong>the</strong> protection <strong>of</strong> breed<strong>in</strong>g grounds from <strong>the</strong> impact <strong>of</strong> tourists<br />

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