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IPM IN PECAN PRODUCTION<br />
143<br />
1989). The basis for this specificity is not fully known, but it occurs sometime later<br />
than 4 days after inoculation during the stage of subcuticular growth (Bracewell &<br />
Stevenson, 1999). Intercropping a mixture of cultivars has been shown to<br />
successfully reduce epidemics of foliar pathogens in other crops (Mundt, 2002), <strong>and</strong><br />
computer simulations based on apple scab show disease reductions of 65–79% after<br />
six generations when planting three cultivars versus a single cultivar (Gessler &<br />
Blaise, 1994). While utilizing that model to guide cultivar placement in new<br />
orchards has been discussed among pecan growers <strong>and</strong> researchers, the benefits of<br />
planting cultivars in blocks for other management purposes usually outweigh the<br />
potential benefits related to scab control.<br />
The race structure of pecan scab populations <strong>and</strong> their ability to adapt to new<br />
cultivars has raised questions regarding the objectives of pecan breeding programs<br />
which involve selecting for scab resistance. Breeders have selected for scab resistance<br />
for years, <strong>and</strong> there have also been attempts to introduce new resistance factors from<br />
other members of the hickory family via interspecific crosses (Graves & Diehl, 1991).<br />
It has generally been recognized that there is a “grace period” after the introduction of<br />
a new cultivar before scab becomes adapted to it, <strong>and</strong> the length of this grace period<br />
varies considerably among cultivars. One objective of the Ge<strong>org</strong>ia breeding program is<br />
to introduce new cultivars with high quality nuts with different resistance genes that<br />
can be transitioned into orchards to maintain diversity (Conner, 2003). The current<br />
Ge<strong>org</strong>ia program is also utilizing DNA markers for resistance genes <strong>and</strong> examining<br />
the physiological basis for scab resistance. Hopefully these efforts will lead to more<br />
durable field resistance in new pecan cultivars.<br />
2.3. Weed Management in Pecan Orchards<br />
Weeds are present throughout the world <strong>and</strong> many interfere with the optimum<br />
production of food <strong>and</strong> fiber. Holm, Doll, Holm, <strong>and</strong> Pancho (1997) state that<br />
approximately 200 plant species world wide are found to cause widespread<br />
problems in our crops. Many of these weeds occur in fruit <strong>and</strong> nut crops. Pecans are<br />
grown in the southern US <strong>and</strong> Mexico, an area that accounts for over 98% of the<br />
world’s pecan production (Pena, 2006). Vines, including annual vines such as the<br />
morningglories (Ipomoea species) <strong>and</strong> perennial vines such as trumpet creeper<br />
(Campsis radicans), <strong>and</strong> both annual <strong>and</strong> perennial grasses such as crabgrass<br />
(Digitaria spp.), goosegrass (Eleusine indica), bahiagrass (Paspalum notatum) <strong>and</strong><br />
bermudagrass (Cynodon dactylon), are some of the most common <strong>and</strong> troublesome<br />
weeds in pecans (Southern Weed Science Society, 2006). Other species commonly<br />
found in pecan orchards include nutsedge (Cyperus species), horsenettle (Solanum<br />
species), arrowleaf sida (Sida rhombifolia), spurges (Euphorbia species), horseweed<br />
(Conyza canadensis), <strong>and</strong> pigweed (Amaranthus species) Colour pictures <strong>and</strong><br />
descriptions are found in the Weed Identification Guide published <strong>by</strong> the Southern<br />
Weed Science Society (2007).<br />
Numerous weed-competition studies have shown the adverse effects of weeds<br />
on agronomic, vegetable, <strong>and</strong> fruit crops (Zimdahl, 2004). These studies generally<br />
document the effects of individual weed species on the growth <strong>and</strong>/or yield of a