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IPM IN PECAN PRODUCTION<br />

143<br />

1989). The basis for this specificity is not fully known, but it occurs sometime later<br />

than 4 days after inoculation during the stage of subcuticular growth (Bracewell &<br />

Stevenson, 1999). Intercropping a mixture of cultivars has been shown to<br />

successfully reduce epidemics of foliar pathogens in other crops (Mundt, 2002), <strong>and</strong><br />

computer simulations based on apple scab show disease reductions of 65–79% after<br />

six generations when planting three cultivars versus a single cultivar (Gessler &<br />

Blaise, 1994). While utilizing that model to guide cultivar placement in new<br />

orchards has been discussed among pecan growers <strong>and</strong> researchers, the benefits of<br />

planting cultivars in blocks for other management purposes usually outweigh the<br />

potential benefits related to scab control.<br />

The race structure of pecan scab populations <strong>and</strong> their ability to adapt to new<br />

cultivars has raised questions regarding the objectives of pecan breeding programs<br />

which involve selecting for scab resistance. Breeders have selected for scab resistance<br />

for years, <strong>and</strong> there have also been attempts to introduce new resistance factors from<br />

other members of the hickory family via interspecific crosses (Graves & Diehl, 1991).<br />

It has generally been recognized that there is a “grace period” after the introduction of<br />

a new cultivar before scab becomes adapted to it, <strong>and</strong> the length of this grace period<br />

varies considerably among cultivars. One objective of the Ge<strong>org</strong>ia breeding program is<br />

to introduce new cultivars with high quality nuts with different resistance genes that<br />

can be transitioned into orchards to maintain diversity (Conner, 2003). The current<br />

Ge<strong>org</strong>ia program is also utilizing DNA markers for resistance genes <strong>and</strong> examining<br />

the physiological basis for scab resistance. Hopefully these efforts will lead to more<br />

durable field resistance in new pecan cultivars.<br />

2.3. Weed Management in Pecan Orchards<br />

Weeds are present throughout the world <strong>and</strong> many interfere with the optimum<br />

production of food <strong>and</strong> fiber. Holm, Doll, Holm, <strong>and</strong> Pancho (1997) state that<br />

approximately 200 plant species world wide are found to cause widespread<br />

problems in our crops. Many of these weeds occur in fruit <strong>and</strong> nut crops. Pecans are<br />

grown in the southern US <strong>and</strong> Mexico, an area that accounts for over 98% of the<br />

world’s pecan production (Pena, 2006). Vines, including annual vines such as the<br />

morningglories (Ipomoea species) <strong>and</strong> perennial vines such as trumpet creeper<br />

(Campsis radicans), <strong>and</strong> both annual <strong>and</strong> perennial grasses such as crabgrass<br />

(Digitaria spp.), goosegrass (Eleusine indica), bahiagrass (Paspalum notatum) <strong>and</strong><br />

bermudagrass (Cynodon dactylon), are some of the most common <strong>and</strong> troublesome<br />

weeds in pecans (Southern Weed Science Society, 2006). Other species commonly<br />

found in pecan orchards include nutsedge (Cyperus species), horsenettle (Solanum<br />

species), arrowleaf sida (Sida rhombifolia), spurges (Euphorbia species), horseweed<br />

(Conyza canadensis), <strong>and</strong> pigweed (Amaranthus species) Colour pictures <strong>and</strong><br />

descriptions are found in the Weed Identification Guide published <strong>by</strong> the Southern<br />

Weed Science Society (2007).<br />

Numerous weed-competition studies have shown the adverse effects of weeds<br />

on agronomic, vegetable, <strong>and</strong> fruit crops (Zimdahl, 2004). These studies generally<br />

document the effects of individual weed species on the growth <strong>and</strong>/or yield of a

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