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214<br />

M.S.T. ABBAS<br />

3.2.3. Locomotion<br />

Flight of the weevils is restricted to day time <strong>and</strong> they have never been observed<br />

as attracted <strong>by</strong> light traps (Nirula, 1956). Al-Khatri <strong>and</strong> Abd-Allah (2003)<br />

studied the daily active periods of R. ferrugineus in date palm plantations in<br />

Oman, using the pheromone traps. They found that the weevils had two periods<br />

of activity, the first between 6 <strong>and</strong> 9 am <strong>and</strong> the second between 6 <strong>and</strong> 9 pm<br />

(sunrise in that area during the period of the study was at 6:15 am <strong>and</strong> sunset<br />

was at 5:50 pm). The weevils were noticed to have no activity during the periods<br />

between 9 am <strong>and</strong> 3 pm <strong>and</strong> between midnight to 3 am. Similarly, Gunawardena<br />

<strong>and</strong> B<strong>and</strong>arage (1995) reported that the periods of RPW activity in Sri Lanka<br />

were between 6 <strong>and</strong> 8 am <strong>and</strong> between 6 <strong>and</strong> 8 pm.<br />

RPW adults are capable of flying as well as crawling. When they are thrown<br />

in the air they fly away in a circle with a buzzing noise <strong>and</strong> quickly disappear,<br />

sometimes l<strong>and</strong>ing shortly <strong>and</strong> boring into the soil. It is believed that the insect<br />

is capable of either flying to a long distance or being carried <strong>by</strong> the winds. It was<br />

noticed that when a weevil was attracted to a pheromone trap it l<strong>and</strong>ed at less<br />

than 1 m apart from the trap, then crawled until reaching it (personal<br />

observation). Chinchilla, Oehlschlager, <strong>and</strong> Gonzaler (1993) reported that the<br />

average flight of R. ferrugineus adults was around 500 m per day <strong>and</strong> a small<br />

portion of the insect migrated up to 1 km per day. Abbas, Hanounik, Shahdad,<br />

<strong>and</strong> Al-Bogham (2006) found that when marked RPWs were released in date<br />

palm plantations, some of them were captured <strong>by</strong> pheromone traps in other<br />

plantations, 1–7 km apart from the release area. Most of such weevils were<br />

captured 3–5 days post release.<br />

4. NATURAL ENEMIES<br />

There are few records about the occurrence of natural enemies of R. ferrugineus,<br />

which might be attributed to the cryptic habitat of the eggs, larvae <strong>and</strong> pupae<br />

which protects them from such natural enemies.<br />

4.1. Parasitoids<br />

Scolia erratica (Hym.: Scoliidae) was reported as an ectoparasitoid of R. ferrugineus<br />

larvae (Nirula, 1956). However, no biological studies on this parasitoid have<br />

been reported. In Brazil, Moura, Mariau, <strong>and</strong> Delabie (1993) <strong>and</strong> Moura,<br />

Resende, <strong>and</strong> Vilela (1995) reported the tachinid Paratheresia menezesi as a<br />

larval-pupal parasitoid of R. palmarum <strong>and</strong> many individuals of the parasitoid<br />

adults emerged from a single pupa. However, attempts were carried out to rear<br />

this parasitoid on R. ferrugineus in the laboratory, but the results were<br />

unsuccessful (unpublished). Nirula (1956) mentioned that both RPW pupae <strong>and</strong><br />

adults were attacked <strong>by</strong> an unknown species of parasitic mite which killed the<br />

pupae <strong>and</strong> reduced the longevity of adults.

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