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Invasive alien species in Switzerland - Schweizer ...

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2 > Vertebrates – Vertebrata 39<br />

urban areas, the normal habitat <strong>in</strong> Central Europe, the environmental impact is probably<br />

less pronounced, although it is an additional omnivorous predator. However, its<br />

impact on structures and foodstuffs and its role as a disease vector <strong>in</strong> human settlements<br />

are unambiguous and the brown rat is controlled <strong>in</strong> many cities.<br />

The raccoon (Procyon lotor (L.)) (see Fact Sheet) is the only member of the family<br />

Procyonidae <strong>in</strong>troduced to Europe. It was <strong>in</strong>troduced from North America and released<br />

<strong>in</strong> Germany <strong>in</strong> 1934. From that and other subsequent releases, it spread through Germany<br />

and reached <strong>Switzerland</strong> <strong>in</strong> 1976. The spread cont<strong>in</strong>ues and it will eventually<br />

<strong>in</strong>habit the lowland parts of <strong>Switzerland</strong> to Genève <strong>in</strong> the south and the Valais. The<br />

impact of the <strong>species</strong> is disputed. A def<strong>in</strong>itive conclusion cannot be reached without<br />

studies be<strong>in</strong>g carried out <strong>in</strong> <strong>Switzerland</strong> to assess its impact on the native wildlife and<br />

as a vector of diseases (Hohmann et al., 2002). This addition to the native fauna may<br />

have to be accepted, s<strong>in</strong>ce control measures are not very effective and its impacts have<br />

not (yet) been demontrated. If <strong>in</strong>cidences of diseases carried by the raccoon <strong>in</strong>crease,<br />

hunters will have to focus more on this <strong>species</strong>.<br />

The only <strong>in</strong>troduced dog (Canidae) <strong>in</strong> Central Europe is the east Asian raccoon dog<br />

(Nyctereutes procyonoides (Gray)) (see Fact Sheet), which was for decades and at<br />

many places <strong>in</strong> the former USSR extensively released as a fur animal. The acclimatization<br />

of the <strong>species</strong> and the value of the fur are questionable. However, it spread westwards<br />

and recently reached France. Incorrect identification based on field observations<br />

by laypeople may be frequent, because they are not familiar with this <strong>species</strong> and it can<br />

be confused with other predators, such as the raccoon. Thus, identification of dead<br />

animals is the most reliable way of evaluat<strong>in</strong>g its spread through Europe. Op<strong>in</strong>ions<br />

about its impact vary between it be<strong>in</strong>g a benign <strong>species</strong> and a <strong>species</strong> caus<strong>in</strong>g severe<br />

impact on native wildlife. One argument is that, as a generalist on abundant food<br />

resources, it does no harm to rare organisms. Others argue that predation on bird<br />

<strong>species</strong> and amphibians might have a negative impact (Kauhala, 1996). The status of<br />

many amphibians as endangered <strong>in</strong> Central Europe and the fact that amphibians were<br />

found <strong>in</strong> proportions of up to 45 % <strong>in</strong> the diet of the raccoon dog (Barbu, 1972) <strong>in</strong>dicates<br />

the potential harm. It also plays a role as a vector of diseases to humans and other<br />

predators (Weidema, 2000). It is recommended that the spread of the raccoon dog<br />

should be monitored and its impact <strong>in</strong> <strong>Switzerland</strong> evaluated. However, a control<br />

programme seems unlikely to achieve success, because of the raccoon dog’s secretive<br />

behaviour and its cont<strong>in</strong>u<strong>in</strong>g <strong>in</strong>vasion from the East. The raccoon dog and the raccoon<br />

are two additional predators with which native animals have to cope. Thus, although<br />

the impacts of the <strong>in</strong>troduced predatory <strong>species</strong> are not so different from those of native<br />

orig<strong>in</strong>, if they add to the total predator load, they are likely <strong>in</strong>crease the predatory<br />

pressure, and destabilize exist<strong>in</strong>g populations and population cycles.<br />

The only established <strong>alien</strong> deer (Cervidae) <strong>in</strong> <strong>Switzerland</strong> is the sika deer (Cervus<br />

nippon Temm<strong>in</strong>ck). It is a fairly small deer <strong>species</strong>, <strong>in</strong>termediate <strong>in</strong> size between the<br />

native roe deer (Capreolus capreolus (L.)) and red deer (Cervus elaphus L.). The<br />

summer coat is chestnut brown to buff brown with white spots, while it is uniform<br />

grey-brown dur<strong>in</strong>g w<strong>in</strong>ter. The rump patch is white with a black upper border and the<br />

white tail has a black stripe. The typical antlers have four po<strong>in</strong>ts each. The sika deer is

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