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Invasive alien species in Switzerland - Schweizer ...

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4 > Insects – Insecta 83<br />

4.8 Dictyoptera<br />

At least five exotic cockroach <strong>species</strong> are known to be established <strong>in</strong> <strong>Switzerland</strong>.<br />

Others may be <strong>in</strong>troduced occasionally with imported goods. Unlike the native cockroaches,<br />

the exotic <strong>species</strong> are synanthropic, i.e. they are always found <strong>in</strong> association<br />

with humans. These cockroaches are of tropical and subtropical orig<strong>in</strong> but they now<br />

occur worldwide. Some <strong>species</strong>, such as Blatta orientalis and Periplaneta americana,<br />

have been cosmopolitan for such a long time that their orig<strong>in</strong> is uncerta<strong>in</strong>. B. orientalis,<br />

P. americana, P. australasiae and B. germanica have been established <strong>in</strong> <strong>Switzerland</strong><br />

for a long time, whereas Supella longipalpa is a recent <strong>in</strong>troduction.<br />

All these <strong>species</strong> are considered as serious urban pests. They are gregarious, nocturnal,<br />

and difficult to eradicate <strong>in</strong> houses. They can eat almost anyth<strong>in</strong>g, e.g. human food and<br />

animal feed, bookb<strong>in</strong>d<strong>in</strong>gs, wallpaper, excrement, leather products, etc. They can carry<br />

organisms caus<strong>in</strong>g human diseases, have a repulsive odour, and can cause allergic<br />

reaction and anxiety <strong>in</strong> some <strong>in</strong>dividuals. In <strong>Switzerland</strong>, the most problematic <strong>species</strong><br />

are B. germanica, S. longipalpa and B. orientalis, whereas Periplaneta spp. are of<br />

m<strong>in</strong>or importance (Landau et al., 1999).<br />

4.9 Isoptera<br />

There is no record of establishment of exotic termites <strong>in</strong> <strong>Switzerland</strong>. However, <strong>in</strong><br />

recent years, subterranean termites of the genus Reticulitermes have <strong>in</strong>vaded new areas<br />

(see Fact Sheet). The North American R. flavipes (Kollar) is <strong>in</strong>troduced <strong>in</strong> some German<br />

and Austrian cities. The southern European R. grassei Clément occurs now <strong>in</strong><br />

southern England and R. santonensis de Feytaud, a <strong>species</strong> of uncerta<strong>in</strong> orig<strong>in</strong> (and<br />

perhaps synonymous with R. flavipes) has spread from south-western to northern<br />

France. These termites undoubtedly have the potential to <strong>in</strong>vade the warmest areas <strong>in</strong><br />

<strong>Switzerland</strong>. They live <strong>in</strong> colonies <strong>in</strong> the soil, ma<strong>in</strong>ly <strong>in</strong> urban areas. They are particularly<br />

harmful to wooden elements <strong>in</strong> build<strong>in</strong>g but can also attack liv<strong>in</strong>g trees. The<br />

spread of subterranean termites <strong>in</strong> France is caus<strong>in</strong>g major concern. New regulations<br />

have been set up to limit the spread.<br />

4.10 Thysanoptera<br />

Many exotic thrips <strong>species</strong> are found, regularly or occasionally, <strong>in</strong> <strong>Switzerland</strong> <strong>in</strong><br />

greenhouses. However, it is not always clear whether they are firmly established <strong>in</strong> the<br />

country. At least four <strong>species</strong> are considered permanently established, ma<strong>in</strong>ly <strong>in</strong> greenhouses.<br />

Three of them, Frankl<strong>in</strong>iella <strong>in</strong>tonsa, F. occidentalis and Heliothrips haemorrhoidalis<br />

are polyphagous, cosmopolitan pests on ornamentals, vegetables, and fruits.<br />

F. occidentalis, the western flower thrips, is among the most destructive greenhouse<br />

pests <strong>in</strong> <strong>Switzerland</strong> (Ebener et al., 1989; Schmidt and Frey, 1995). It is also a vector of<br />

various viruses. The pest status of F. occidentalis is relatively recent. It occurs naturally<br />

<strong>in</strong> North America on wild flowers and emerged as a greenhouse pest <strong>in</strong> the Neth-

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