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Leiodidae or small carrion beetles, are more elliptical in body shape and feed on carrion or dung. Some species are specializecl occupants of ant nests, caves (but not in Canada), rr the nests and burrows of vertebrates. Members irf the Leptininae family (= Leptinidae), or mammal rrest beetles and beaver beetles, are nest scavengers and ectoparasites on small mammal hosts or on beavers. Collec;ing Leiodidae usually requires a concentration on the particular habits or habitats of the genera or species sought. They can be collected in the following ways: in baited pitfall traps; by sifting and Berlese extraction of litter or fungi; by sweeping forest vegc,tation on still, warm, and humid evenings; and by use,, of large-area flight intercept traps. The leiodid fauna of Canada and Alaska is only partly known. Several genera have been reviewed and revised recently, and others are being prepared. No single publication treats the entire fauna, but a generic review of the North American fauna is available (Peck 1990). The largest faunal works (Hatch 1933, 1957; Jeannel 1936) are no longer completely reliable. Distributions and bionomics of the species are usually incompletely known, and many species in the northern United States may eventually be found in southern Canada. AK (36);YK (18); NT (16); BC (84);AB (39)t SK (18); MB (26); oN (57); PQ (53); NB (1e);NS (15); t B (7); NF(10) . H(6);I(1) Subfamily LEIODINAE Tribe Hydnobiini Genus HYDIIOBIUS Schmidt H. contortus Hatch H. crestonensis Hatch H. femoratus Hatch H. longulus l,eConte longidets LeConte H. luggeri H rtch H. matthewsii Crotch H. obtusus L,eConte H. pumilus L,eConte hiseri Hatch latidens LeConte H. simulator W.J. Brown H. stacesmit ri Hatch H. strigilatu r Horn H. substriatrLs LeOonte curuid.ens LeConte H. validus \[.J. Brown AK --ec-- -BC _BCAB BC _BC _BC _BC _BC AB SK ON PQ pe PQ NS Genus TRIAIRTHRON Markel T. lecontei Horn cedonuli'.i Schaufuss _BC Genus ANO(]DUS LeConte Neocyrtlsa W.J. Brown Key to species: Daffner (1988) A. fusciclavus (Fall t sternitu; (Hatch) A. insolitus tW.J. Brownt A. obsoletus Melsheimer) blandis :; ima (Zimmermann) A. potens tW.J. Brownt A. puritanus {Fallt A. secretus (W.J. Brown) A. superans Fall) Tribe Leiodini BC BC BC SK oN-- _ONPQ _ONPQ -oNPQ AB_MBON Genus LIONOTHUS W.J. Brown Key to No;th American species: Daffner (1988) L. forticornis Daffner L. ulkei W.J Brown -oN -oN PQ 78
Leiodidae Genus CYRTUSA Erichson Caenocyrta W.J. Brown Key to North American species: Daffner (1988) C. subtestacea (Gyllenhal)* picipennis (LeConte) BC oN PQ Genus LIOCYRTUSA Daffner Key to North American species: Daffner (1988) L. luggeri (Hatch) L. nigriclavis (Hlisnikovsky)* Genus ECARINOSPHAERULA Hatch E. ecarinata (Horn) Genus ISOPLASTUS Horn I. fossor Horn uncus Wheeler _BCABSKMBONPQNB _BC_ONPQ BC oN PQ Genus LEIODES Latreille L. assimilis (LeConte) L. bradorata W.J. Brown L. canadensis (W.J. Brown) L. collaris (LeConte) L. conferta (LeConte) L. curvata (Mannerheim) morula LeConte L. diffrcilis (Horn) L. dilatata Hatch L. grossa Hatch L. impersonata W.J. Brown L. lateritia (Mannerheim) L. lostine (Hatch) L. merkeliana (Horn) L. mirabilis (Hatch) L. paludicola (Crotch) L. puncticollis (C.G. Thomson) L. punctostriata Kirby indistincta (LeConte) laefo (Mannerheim) punctatostriato Horn punctistriata LeConte L. punctulata (GyIlenhal) L. rhaetica (Erichson) L. rufrpes (Gebler) L. serripes Hatch L. strigata (LeConte) L. tenuis Hatch L. triepkei (Schmidt) L. valida (Horn) yyNrBC AKYK_BC AK BC AB AB AB - - - ncy -BCAB AKBC_ _BCAB NT BC _BC -BC AKYK_BCAB AK YK NT BC AB SK SK SK MB ON PQ PQ MB MB ON PQ _ 9*PQ -oNpe nfn ON Pq MB ON PQ AKYKNTBCAB_MBONPQ AK AK YK NT BC AB SK MB ON _BCABSK AKBCABSK-ONPQ _BC -YKNT-AB-MBONPQ NTBCAB_MBONPQ NB NB NB NB NB NB NB NS NS NS LB NF LB NF LB NF Genus ZEADOLOPUS Broun Apheloplastus W.J. Brown Key to North American species: Daffer (1988) Z. egenus (LeConte) impubis (Zimmermann) oN PQ Genus AGLYPTINUS Cockerell A. laevis (LeConte) Tlibe Scotocryptini oN PQ Genus COLENIS Erichson C. impunctata LeConte Tribe Pseudoliodini BC MB ON PQ NB NS 79
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l*l 331:!y" Checklist of of Canada
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oMinister of Supply anrl Services C
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Family Cerylonidae (J.M. Campbell)
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SUMMARY All species and subspecies
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INTRODUCTION Beetles represent the
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Micromalthidae Order Coleoptera Sub
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Carabidae Family CARABIDAE ground b
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Carabidae C. tranqueba'ica borealis
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Carabidae lapilayi Castelnau lecont
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Carabidae Genus NEBRIA Latreille N.
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Carabidae Genus ELAI'HRUS Fabricius
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Carabidae D. nigricornis Motschulsk
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Carabidae T. apicalis Motschulskyx
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Carabidae atnicum Casey deceptor Ca
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Carabidae B. nevadense Illke uiatic
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Carabidae B. dorsale Say B, graphic
Page 36 and 37: Carabidae B. connivens (LeConte) di
Page 38 and 39: Carabidae P. foveocollis (Eschschol
Page 40 and 41: Carabidae P. ecarinatur, Hatch P. h
Page 42 and 43: Carabidae P. Ieconteianus (Lutshnik
Page 44 and 45: Carabidae adstricta (Putzeys) caten
Page 46 and 47: Carabidae ferruginea Casey liquida
Page 48 and 49: Carabidae litoreus Jasey pensylucin
Page 50 and 51: Carabidae foueicollis LeConte mutab
Page 52 and 53: Carabidae A. viridescenr LeConte ch
Page 54 and 55: Carabidae Genus TRIC JOCELLUS Gangl
Page 56 and 57: Carabidae curlipet nis Casey expert
Page 58 and 59: Carabidae C. pusillus Say elegant,i
Page 60 and 61: Carabidae O. micans LcConte O. parm
Page 62 and 63: Carabidae A. cupripenne (Say) NT BC
Page 64 and 65: Carabidae pacatus ( Casey) sinuatus
Page 66 and 67: Carabidae truckeens s Casey uermicu
Page 68 and 69: Haliplidae G. janus (Fablicius) aha
Page 70 and 71: Dytiscidae be omnivorous. A detaile
Page 72 and 73: Dytiscidae macularis (LeOonte) micr
Page 74 and 75: Dy'tiscidae H. dentellus Fall AK YK
Page 76 and 77: Dytiscidae A. aeruginosrs Aub6 disp
Page 78 and 79: Dytiscidae C. densus LeConte AK YK
Page 80 and 81: Gytininae G. liberus (Sr,Ly) brunni
Page 82 and 83: Hydraenidae Suborder Polyphaga Supe
Page 84 and 85: Limuiodidae Subfamily ACROTRICHIINA
Page 88 and 89: Leiodidae Tribe Agathidini Genus AN
Page 90 and 91: Scydmaenidr,Le Subfamily LEPTININAE
Page 92 and 93: Silphidae S. retrusus (Casey) S. zi
Page 94 and 95: Staphylinidae infodiens Mannerheim
Page 96 and 97: Staphylinida I E. californicum (Fau
Page 98 and 99: Staphylinidae angulare Fanvel colum
Page 100 and 101: Staphylinidar,, Subfamily OLISTHAER
Page 102 and 103: Staphylinidr,Le B. opaculus LeConte
Page 104 and 105: Staphylinidu e Genus TACFIINUS Grav
Page 106 and 107: Staphylinidr.e Tlibe Bolitobiini Ge
Page 108 and 109: Staphylinida e O. leechi Lohse O. m
Page 110 and 111: Staphylinidae Genus LIOGLUTA C.G. T
Page 112 and 113: Staphylinidae Genus AJ-ECiCHARA Gra
Page 114 and 115: Staphylinidae globosa l{elsheimer i
Page 116 and 117: Staphyiinida: Genus LIPAIiOCEPHALUS
Page 118 and 119: Staphylinidae S. pollens (Casey) pa
Page 120 and 121: Staphylinid re S. pusillimus Puthz
Page 122 and 123: Staphylinidr.e (Subgenus PLATYMEDON
Page 124 and 125: Staphylinidr:e Genus LOBIiATHIUM Mu
Page 126 and 127: Staphylinidar,r P. concinnus t Grav
Page 128 and 129: Staphylinide:e Tribe Staphylinini G
Page 130 and 131: Staphyiinida: laeuigati ts (Gyllenh
Page 132 and 133: Scaphidiidae Genus STIC'IOLINUS Cas
Page 134 and 135: Pselaphidae Genus MEGARAFONUS Casey
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Pselaphidae Genus MAC HAXRODES Bren
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Hydrophilicae Superfamily Hydrophil
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Hydrophili
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Hydrophilidae Genus CERCYON Leach (
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Histeridae Genus PLEGADERUS Erichso
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Histeridae Genus NEOI'ACHYLOPUS Rei
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Histeridae H. defectus .f.E. LeCont
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Scirtidae C. pubescens Redtenbacher
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Lucanidae Subfamilv SYNDESINAE Trib
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Scarabaeidae Genus CAIITHON Hoffman
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Scarabaeic ae A. erraticus (Linn6)t
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Scarabaeidae Genus BOL,IOCEROSOMA S
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Scarabaeid,re D. tristis K rby MB O
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Scarabaeida': D. vicina (Fall) colu
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Scarabaeidae Tribe Pentodontinr Gen
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Byrrhidae Genus TRITIHIOTINUS Casey
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Buprestida: Tlibe Pedilophorini Gen
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Buprestidae Genus SPE CTRAJ-IA Case
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Buprestidae sexmaculata Hausmann se
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Buprestidae Subfamily CHRYSOBOTHRIN
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Buprestidar: desertus LeConte plumb
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Heteroceridae Genus MICIIOAUGYLES P
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Elmidae H. basalis L,:Conte H. colu
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Artematopidae Family PSEPHENIDAE wa
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Elateridae Genus DR.A.STERIUS Eschs
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Elateridae D. denticor nis { Kirby
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Elateridae C. moerens. tLeConte) C.
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Elateridae Genus SERICUS Eschcholtz
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Elateridae A. sayi (Let)onter - ON
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Eucnemidae: FamilyTHROSCIDAE throsc
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Lycidae Genus EPIPHANIS Eschscholtz
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Lampyrrida e FamiIyPHENGODIDAE glow
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Cantharid,re Family CANTHARIDAE sol
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Cantharidaer R. imbecillis (LeConte
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Dermestida,: Genus PELi|ASTICA Mann
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Dermestidae T. sinistrurn FaIl AK T
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Bostrichidac Genus PSOI, Herbst Acr
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Anobiidae Subfamily ERNOBIINAE Trib
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Anobiidae Genus EIIIRILLETTA Fall O
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Trogossitida e Genus PSEI.IDEUROSTU
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Cleridae Genus TEI'IEBROIDES Piller
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Cleridae E. nigrifrons tSayt * ON P
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Melyridae M. floricola Marshall - B
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Nitidulidaer Family NITIDULIDAE sap
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Nitidulidae Genus OMOSITA Erichson
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Rhizophagi lae Family RHIZOPIIAGIDA
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Cucujidae Genus CRY FIOLESTES Gangl
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Cryptophag idae C. bidentatus Makli
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Languriidze Subsequi:ntly (1968), h
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Phalacrida,r cincta Lacordafte dimi
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Corylophidae Family CORYLOPHIDAE mi
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Coccinellidae Genus SCYMNUS Kugelan
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Coccinellidae schuhi }{atch simuloi
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Coccinellidz,.e Genus HIPI'ODAMIA D
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Coccinellidi,'e C. alta W.J. Brown
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Endomychidae Subfamily MYCETAEINAE
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Lathridiidae L. hirtus Gyllenhalf L
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Mycetophagidae baker;'.Barber breui
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Ciidae Family CIIDAE minute tree-fu
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Melandryidre Subfamily IIALLOMENINA
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Mordellida: Genus SEF,ROPAIPUS Hell
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Rhipiphoridae M. ornata (trfelsheim
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Zopheridae Genus COLYDIUM Fabricrus
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Tenebrionldae punchtta Casey regulc
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Tenebrionidae P. americanum Casteln
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Tenebrionidae T?ibe Helopini Genus
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Tenebrionilae P. pectinat.r Hopping
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Cephaloidae Subfamily GONIADERINAE
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Meloidae M. niger Kirby hirbyi Drll
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Oedemeridae Z. sayi Wickham imrnact
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Pythidae Family BORIDAE J.M. Campbe
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Salpingidae Family SALPINGIDAE narr
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Anthicidaer heluin.us Casey minuscu
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Pedilidae Family PEDILIDAE pedilid
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Scraptiidac Subfamily ANASPIDINAE T
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Cerambycidee ambiguus Casey angustu
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Cerambycii.ae T. schwarzilnum Casey
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Cerambycide.e Tlibe Hyboderini Genu
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Cerambycidae grandi; Casey obscur,t
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Cerambycirlae C. margini
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CerambyciJae Genus ULOCIIAETES LeOo
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Cerambycidae Genus CO IITODERA Muls
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Cerambycidae T. zebra (Olivier) aur
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Cerambycidae B. dehiscens (LeConte)
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Cerambycirlae Genus LOI'HOPOGONIUS
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Cerambycidae Genus URGLEPTES Dillon
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Bruchidae T. femoratus LeConte amni
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Chrysome lidae Genus PL,A.TEUMARIS
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Chrysomelidae Genus ZEIIGOPHORA Kun
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Chrysomelidae P. obsoletr.,s Suffri
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Chrysomel:idae S. hamata Horn - BC
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Chrysomelirlae Genus TIMARCHA Latre
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Chrysomelirlae C. hudsonica W.J. Br
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Chrysomelic;.ae Subfamily GALERUCIN
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ChrysomeliCae Genus PSELIDOLUPERUS
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ChrysomeLidae A. bimargirrata Say p
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Chrysomel,dae parua Blatch\ey pensy
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ChrysomelLdae Genus STEiNOPODIUS Ho
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Anthribidae Genus CIMBERIS des Gozi
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Rhynchitidrre FamilyATTELABIDAE lea
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Apionidae Family APIONIDAE pear-sha
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Curculionidae Tribe Barynotini Genu
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Curculionidae Genus AGRONUS Horn A.
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Curculionidae (Subgenus IIIOMIDUS C
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Curculioni lae Genus HYPERA Germar
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Curculioni*1ae H. congener Dalla To
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Curculioniclae Genus SMICRONYX Scho
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Curculionitiae M. myrmex (Herbst) a
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Curculionidae A. nigrinus.Boheman A
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Curculionid:re L. bischoffi tBlatch
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Curculionidae T. capitale \Vibmer T
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Curculionidae C. oregonersis Dietz
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Curculionidae demissa Casey dolosa
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Curcuiionii.ae Genus SPHENOPHORUS S
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Scolytidae Genus RHIINCOLUS Germar
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Scolytidae D. murrayar:.ae Hopkins
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Scolytidae Tribe Ipini Genus PITYOG
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Scolytidae Genus XYLI.BORUS Eichhof
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Scolytidae P. confinis LeConte BC P
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Halictophagidae Order Strepsiptera
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REFERENCES Abdullah, M. 1965. World
Page 378 and 379:
Beal, R.S., Jr. 1985b. Review of Ne
Page 380 and 381:
Bousquet, Y. 1990. A review of the
Page 382 and 383:
Campbell, J.M. 1978b. A revision of
Page 384 and 385:
Colonnelli, E. 1979. Note su Panoph
Page 386 and 387:
Ford, E.J. 1973. A revision of the
Page 388 and 389:
Hacker, II.A. 1968. The species of
Page 390 and 391:
Hinton, H.E. 1948. A synopsis of th
Page 392 and 393:
Klimaszewski, J. 1982. Studies of M
Page 394 and 395:
Linsley, E.G. 1964. The Cerambycida
Page 396 and 397:
Neboiss, A. 1984. Reclassification
Page 398 and 399:
Reyes-Castillo, P. 1970. Coleoptera
Page 400 and 401:
Smetana, A. 1978. Revision of the s
Page 402 and 403:
Wallis, J.B. 1973. An annotated lis
Page 404 and 405:
Young, I).K. 1975. A revision of th
Page 406 and 407:
Agronus Horn 332 Agyrtecanus Reitte
Page 408 and 409:
Aseminae 279 Asemini 279 Asemobius
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Callisten,',a Vacher de Lapouge 11
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Coelini 254 Coptotomini 70 Coelocep
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Dasy'bes Faykull 212 Dasytinae 212
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Endomychidae 237 Endomychinae 239 E
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Gibbiinae 205 Gibbium Scopoli 205 G
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Hydrobiinae 131 Hydrobiini 131 Hydr
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Leptotrachelus Latreille 56 Leptotr
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Meligethes Stephens 217 Meligethina
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Neophytobius Wagner 351 Neopolya.rt
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Pachyrhjnus Schcinherr SB0 Pachy rh
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Physorhini 184 Phytobiini 350 Phyto
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Pseudota Casey 101 Pseudothysanoes
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Scotocryptini 79 Scraptia Latreille
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Tachyporu.s Gravenhorst 97 Tachypte
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Uloma Ltrtreille 257 Ulomini 257 Un
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