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Silphidae S. retrusus (Casey) S. zimmermz:.nni Schaum punctatus Casey oll PQ Family MICROPEPLIDAE micropeplid beetles J.M. Campbell Micropeplidae is a small family of beetles with 10 species known in Canada and Alaska. They are found in a variety of habitats, including deciduous and conifr,,rous leaf litter, inside beaver lodges, in Carex clurnps growing in shallow water, on freshly cut stumps of fir and cedar and possibly associated with the fungus Pyrobema, which is found in forests after a firr:. Matthews (1970) described two species, M. hoogervdorni and M. hopkinsi, from Pliocene deposits irr western Alaska. Little is known about the feeding habits of the species; however, both larvae and adults are assumed to feed ott fungal spores. The family is well known in North .t\merica. Campbell (1968) revised the New World spr;cies and published a catalog of the North Americatr species (1984c). AK (a); YK (1); NT (1); BC (8); AB (a); SK. (2); MB (1); ON (5);PQ (a) ' H(1) Tribe Micropeplini Genus KALI|jSUS LeConte K. nitidus Lc Conte BC Genus MICROPEPLUS Latreille Anhenooeplus Koch Key to North American species: Campbell (1973b) M. browni Cr'Lmpbell M. brunneus Maklin M. cribratus LeConte M. laticollis li{eklin M. punctatus LeConte costatus Maklin oregonensis Hatch' M. robustus ()ampbell M. sculptus l,eConte obliquus LeConte M. smetanai Campbell M. tesserula Curtis* costatus LeConte costipenzis Makiin AK AK AK AK YK NT BC BC BC BC BC BC BC AB AB AB AB SK sx -oNPQ -oN _ONPQ ON PQ MB ON PQ Family SILPHIDAE carrion beetles J.M. Campbell The ca.rrion beetles belong to a family of about 175 species found primarily in the Northern Hemisphere, with a few species having ranges extending into South America, southeast Asia, and Australia. Twenty-five species are known to occur in Canada and Alaska. Carrion beetles are common, and because oftheir large size and often bright colors they are frequently collected. Both adults and Iarvae ofmost species feed on carcasses ofvertebrates; other species are scavengers, and some species of Aclypea are phy'tophagous and may be considered to be pests ofcultivated crops such as spinach. The family name Silphidae is userl in the restricted sense of Lawrence (I982a), Lawrence and Newton (1982), and Anderson and Peck (198{5). These authors exclude the agyrtids, treating them es a separate family, Agyrtidae. The Canadian and Alaskan species are well known taxonomically, based on a handbook ofthe species by Anderson and Peck (1985). AK (8); YK (6); NT (9); BC (13); AB (17); SK (18); MB (18); ON (18); PQ (15); NB (10); NS (11); PE (7); LB (1); NF (6) . H (4) 84
Silphidae Subfamily SILPHINAE Genus NECRODES Leach Proton ec rod es Portevin N. surinamensis (Fabricius) - BC AB SK MB ON PQ NB NS PE - NF Genus THANATOPHILUS Leach Philas Portevin T. coloradensis (Wickham) AK BC obalskii (Portevin) T. lapponicus (Herbst)* AK YK NT BC AB SK MB ON PQ NB NS - LB NF cal ifor ni cus (Mannerheim) caudatus (Say) tuberculatus (Germar) T. sagax (Mannerheim) AK YK NT BC AB SK MB ON PQ T. trituberculatus (Kirby) AK - NT - AB SK MB ON Genus OICEOPTOMA Leach O. inaequale tFabricius) - ON PQ O. noveboracense (Forster) AB SK MB ON PQ NB NS PE marginale (Fabricius) marginata Kirby Genus NECROPHILA Kirby & Spence N. americana {Linn6} MB ON PQ NB NS affinis (I{ttby) canadensis (Kirby) peltata (Catesby) terminata (KJrby) Genus HETEROSILPHA Portevin H. ramosa {Say) NT BC AB SK MB ON ceruaria (Mannerheim) Genus ACLYPEA Reitter Blitophaga Reitter A. bituberosa {Leconte) NT BC AB SK MB A. opaca tl-innet* AX YK NT Subfamily NICROPHORINAE Genus NICROPHORUS Fabricius Neonicrophorus Hatch N. americanus Olivier grandis Fabricius - ON pe uirginicus Fr6lich N. carolinus (Linn6J med.iatus Fabricius AB mysticallis Angell N. defodiens Mannerheim binotatus Portevin AK YK NT BC AB sK MB oN PQ NB NS pE - NF conuersator Walker hadjakensis Portevin lateralis Porte.rin mannerheimi Portevin plagiatus Motschulsky pollinctor LeConte N. guttula Motschulsky hecate Bland - BC AB SK immaculosis }{atch punctostriatus Pietce N. hybridus Hatch & Angell - BC AB SK MB ON N. investigator Zetterstedt* AK YK NT BC AB SK MB oN Pe - NS pE - NF aleuticus Gistel confossor LeConte 85
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l*l 331:!y" Checklist of of Canada
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oMinister of Supply anrl Services C
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Family Cerylonidae (J.M. Campbell)
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SUMMARY All species and subspecies
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INTRODUCTION Beetles represent the
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Micromalthidae Order Coleoptera Sub
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Carabidae Family CARABIDAE ground b
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Carabidae C. tranqueba'ica borealis
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Carabidae lapilayi Castelnau lecont
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Carabidae Genus NEBRIA Latreille N.
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Carabidae Genus ELAI'HRUS Fabricius
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Carabidae D. nigricornis Motschulsk
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Carabidae T. apicalis Motschulskyx
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Carabidae atnicum Casey deceptor Ca
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Carabidae B. nevadense Illke uiatic
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Carabidae B. dorsale Say B, graphic
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Carabidae B. connivens (LeConte) di
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Carabidae P. foveocollis (Eschschol
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Carabidae P. ecarinatur, Hatch P. h
Page 42 and 43: Carabidae P. Ieconteianus (Lutshnik
Page 44 and 45: Carabidae adstricta (Putzeys) caten
Page 46 and 47: Carabidae ferruginea Casey liquida
Page 48 and 49: Carabidae litoreus Jasey pensylucin
Page 50 and 51: Carabidae foueicollis LeConte mutab
Page 52 and 53: Carabidae A. viridescenr LeConte ch
Page 54 and 55: Carabidae Genus TRIC JOCELLUS Gangl
Page 56 and 57: Carabidae curlipet nis Casey expert
Page 58 and 59: Carabidae C. pusillus Say elegant,i
Page 60 and 61: Carabidae O. micans LcConte O. parm
Page 62 and 63: Carabidae A. cupripenne (Say) NT BC
Page 64 and 65: Carabidae pacatus ( Casey) sinuatus
Page 66 and 67: Carabidae truckeens s Casey uermicu
Page 68 and 69: Haliplidae G. janus (Fablicius) aha
Page 70 and 71: Dytiscidae be omnivorous. A detaile
Page 72 and 73: Dytiscidae macularis (LeOonte) micr
Page 74 and 75: Dy'tiscidae H. dentellus Fall AK YK
Page 76 and 77: Dytiscidae A. aeruginosrs Aub6 disp
Page 78 and 79: Dytiscidae C. densus LeConte AK YK
Page 80 and 81: Gytininae G. liberus (Sr,Ly) brunni
Page 82 and 83: Hydraenidae Suborder Polyphaga Supe
Page 84 and 85: Limuiodidae Subfamily ACROTRICHIINA
Page 86 and 87: Leiodidae or small carrion beetles,
Page 88 and 89: Leiodidae Tribe Agathidini Genus AN
Page 90 and 91: Scydmaenidr,Le Subfamily LEPTININAE
Page 94 and 95: Staphylinidae infodiens Mannerheim
Page 96 and 97: Staphylinida I E. californicum (Fau
Page 98 and 99: Staphylinidae angulare Fanvel colum
Page 100 and 101: Staphylinidar,, Subfamily OLISTHAER
Page 102 and 103: Staphylinidr,Le B. opaculus LeConte
Page 104 and 105: Staphylinidu e Genus TACFIINUS Grav
Page 106 and 107: Staphylinidr.e Tlibe Bolitobiini Ge
Page 108 and 109: Staphylinida e O. leechi Lohse O. m
Page 110 and 111: Staphylinidae Genus LIOGLUTA C.G. T
Page 112 and 113: Staphylinidae Genus AJ-ECiCHARA Gra
Page 114 and 115: Staphylinidae globosa l{elsheimer i
Page 116 and 117: Staphyiinida: Genus LIPAIiOCEPHALUS
Page 118 and 119: Staphylinidae S. pollens (Casey) pa
Page 120 and 121: Staphylinid re S. pusillimus Puthz
Page 122 and 123: Staphylinidr.e (Subgenus PLATYMEDON
Page 124 and 125: Staphylinidr:e Genus LOBIiATHIUM Mu
Page 126 and 127: Staphylinidar,r P. concinnus t Grav
Page 128 and 129: Staphylinide:e Tribe Staphylinini G
Page 130 and 131: Staphyiinida: laeuigati ts (Gyllenh
Page 132 and 133: Scaphidiidae Genus STIC'IOLINUS Cas
Page 134 and 135: Pselaphidae Genus MEGARAFONUS Casey
Page 136 and 137: Pselaphidae Genus MAC HAXRODES Bren
Page 138 and 139: Hydrophilicae Superfamily Hydrophil
Page 140 and 141: Hydrophili
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Hydrophilidae Genus CERCYON Leach (
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Histeridae Genus PLEGADERUS Erichso
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Histeridae Genus NEOI'ACHYLOPUS Rei
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Histeridae H. defectus .f.E. LeCont
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Scirtidae C. pubescens Redtenbacher
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Lucanidae Subfamilv SYNDESINAE Trib
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Scarabaeidae Genus CAIITHON Hoffman
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Scarabaeic ae A. erraticus (Linn6)t
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Scarabaeidae Genus BOL,IOCEROSOMA S
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Scarabaeid,re D. tristis K rby MB O
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Scarabaeida': D. vicina (Fall) colu
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Scarabaeidae Tribe Pentodontinr Gen
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Byrrhidae Genus TRITIHIOTINUS Casey
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Buprestida: Tlibe Pedilophorini Gen
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Buprestidae Genus SPE CTRAJ-IA Case
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Buprestidae sexmaculata Hausmann se
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Buprestidae Subfamily CHRYSOBOTHRIN
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Buprestidar: desertus LeConte plumb
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Heteroceridae Genus MICIIOAUGYLES P
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Elmidae H. basalis L,:Conte H. colu
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Artematopidae Family PSEPHENIDAE wa
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Elateridae Genus DR.A.STERIUS Eschs
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Elateridae D. denticor nis { Kirby
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Elateridae C. moerens. tLeConte) C.
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Elateridae Genus SERICUS Eschcholtz
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Elateridae A. sayi (Let)onter - ON
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Eucnemidae: FamilyTHROSCIDAE throsc
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Lycidae Genus EPIPHANIS Eschscholtz
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Lampyrrida e FamiIyPHENGODIDAE glow
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Cantharid,re Family CANTHARIDAE sol
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Cantharidaer R. imbecillis (LeConte
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Dermestida,: Genus PELi|ASTICA Mann
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Dermestidae T. sinistrurn FaIl AK T
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Bostrichidac Genus PSOI, Herbst Acr
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Anobiidae Subfamily ERNOBIINAE Trib
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Anobiidae Genus EIIIRILLETTA Fall O
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Trogossitida e Genus PSEI.IDEUROSTU
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Cleridae Genus TEI'IEBROIDES Piller
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Cleridae E. nigrifrons tSayt * ON P
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Melyridae M. floricola Marshall - B
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Nitidulidaer Family NITIDULIDAE sap
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Nitidulidae Genus OMOSITA Erichson
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Rhizophagi lae Family RHIZOPIIAGIDA
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Cucujidae Genus CRY FIOLESTES Gangl
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Cryptophag idae C. bidentatus Makli
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Languriidze Subsequi:ntly (1968), h
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Phalacrida,r cincta Lacordafte dimi
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Corylophidae Family CORYLOPHIDAE mi
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Coccinellidae Genus SCYMNUS Kugelan
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Coccinellidae schuhi }{atch simuloi
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Coccinellidz,.e Genus HIPI'ODAMIA D
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Coccinellidi,'e C. alta W.J. Brown
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Endomychidae Subfamily MYCETAEINAE
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Lathridiidae L. hirtus Gyllenhalf L
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Mycetophagidae baker;'.Barber breui
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Ciidae Family CIIDAE minute tree-fu
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Melandryidre Subfamily IIALLOMENINA
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Mordellida: Genus SEF,ROPAIPUS Hell
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Rhipiphoridae M. ornata (trfelsheim
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Zopheridae Genus COLYDIUM Fabricrus
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Tenebrionldae punchtta Casey regulc
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Tenebrionidae P. americanum Casteln
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Tenebrionidae T?ibe Helopini Genus
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Tenebrionilae P. pectinat.r Hopping
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Cephaloidae Subfamily GONIADERINAE
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Meloidae M. niger Kirby hirbyi Drll
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Oedemeridae Z. sayi Wickham imrnact
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Pythidae Family BORIDAE J.M. Campbe
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Salpingidae Family SALPINGIDAE narr
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Anthicidaer heluin.us Casey minuscu
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Pedilidae Family PEDILIDAE pedilid
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Scraptiidac Subfamily ANASPIDINAE T
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Cerambycidee ambiguus Casey angustu
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Cerambycii.ae T. schwarzilnum Casey
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Cerambycide.e Tlibe Hyboderini Genu
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Cerambycidae grandi; Casey obscur,t
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Cerambycirlae C. margini
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CerambyciJae Genus ULOCIIAETES LeOo
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Cerambycidae Genus CO IITODERA Muls
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Cerambycidae T. zebra (Olivier) aur
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Cerambycidae B. dehiscens (LeConte)
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Cerambycirlae Genus LOI'HOPOGONIUS
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Cerambycidae Genus URGLEPTES Dillon
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Bruchidae T. femoratus LeConte amni
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Chrysome lidae Genus PL,A.TEUMARIS
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Chrysomelidae Genus ZEIIGOPHORA Kun
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Chrysomelidae P. obsoletr.,s Suffri
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Chrysomel:idae S. hamata Horn - BC
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Chrysomelirlae Genus TIMARCHA Latre
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Chrysomelirlae C. hudsonica W.J. Br
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Chrysomelic;.ae Subfamily GALERUCIN
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ChrysomeliCae Genus PSELIDOLUPERUS
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ChrysomeLidae A. bimargirrata Say p
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Chrysomel,dae parua Blatch\ey pensy
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ChrysomelLdae Genus STEiNOPODIUS Ho
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Anthribidae Genus CIMBERIS des Gozi
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Rhynchitidrre FamilyATTELABIDAE lea
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Apionidae Family APIONIDAE pear-sha
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Curculionidae Tribe Barynotini Genu
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Curculionidae Genus AGRONUS Horn A.
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Curculionidae (Subgenus IIIOMIDUS C
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Curculioni lae Genus HYPERA Germar
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Curculioni*1ae H. congener Dalla To
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Curculioniclae Genus SMICRONYX Scho
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Curculionitiae M. myrmex (Herbst) a
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Curculionidae A. nigrinus.Boheman A
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Curculionid:re L. bischoffi tBlatch
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Curculionidae T. capitale \Vibmer T
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Curculionidae C. oregonersis Dietz
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Curculionidae demissa Casey dolosa
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Curcuiionii.ae Genus SPHENOPHORUS S
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Scolytidae Genus RHIINCOLUS Germar
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Scolytidae D. murrayar:.ae Hopkins
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Scolytidae Tribe Ipini Genus PITYOG
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Scolytidae Genus XYLI.BORUS Eichhof
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Scolytidae P. confinis LeConte BC P
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Halictophagidae Order Strepsiptera
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REFERENCES Abdullah, M. 1965. World
Page 378 and 379:
Beal, R.S., Jr. 1985b. Review of Ne
Page 380 and 381:
Bousquet, Y. 1990. A review of the
Page 382 and 383:
Campbell, J.M. 1978b. A revision of
Page 384 and 385:
Colonnelli, E. 1979. Note su Panoph
Page 386 and 387:
Ford, E.J. 1973. A revision of the
Page 388 and 389:
Hacker, II.A. 1968. The species of
Page 390 and 391:
Hinton, H.E. 1948. A synopsis of th
Page 392 and 393:
Klimaszewski, J. 1982. Studies of M
Page 394 and 395:
Linsley, E.G. 1964. The Cerambycida
Page 396 and 397:
Neboiss, A. 1984. Reclassification
Page 398 and 399:
Reyes-Castillo, P. 1970. Coleoptera
Page 400 and 401:
Smetana, A. 1978. Revision of the s
Page 402 and 403:
Wallis, J.B. 1973. An annotated lis
Page 404 and 405:
Young, I).K. 1975. A revision of th
Page 406 and 407:
Agronus Horn 332 Agyrtecanus Reitte
Page 408 and 409:
Aseminae 279 Asemini 279 Asemobius
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Callisten,',a Vacher de Lapouge 11
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Coelini 254 Coptotomini 70 Coelocep
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Dasy'bes Faykull 212 Dasytinae 212
Page 416 and 417:
Endomychidae 237 Endomychinae 239 E
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Gibbiinae 205 Gibbium Scopoli 205 G
Page 420 and 421:
Hydrobiinae 131 Hydrobiini 131 Hydr
Page 422 and 423:
Leptotrachelus Latreille 56 Leptotr
Page 424 and 425:
Meligethes Stephens 217 Meligethina
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Neophytobius Wagner 351 Neopolya.rt
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Pachyrhjnus Schcinherr SB0 Pachy rh
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Physorhini 184 Phytobiini 350 Phyto
Page 432 and 433:
Pseudota Casey 101 Pseudothysanoes
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Scotocryptini 79 Scraptia Latreille
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Tachyporu.s Gravenhorst 97 Tachypte
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Uloma Ltrtreille 257 Ulomini 257 Un
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