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TABLE 2. ORIENTATION OF BROWN BEAR DENS ON KODIAK ISLAND AND THE ALASKA PENINSULA. N NE E SE S SW W NW Kodiak Island Females w/yearlings Singles Unidentified 1 1 20 12 8 1 8 10 1 1 7 3 1 6 Total 22 12 8 9 10 9 3 7 Alaska Peninsula Females w/cubs Females w/yearlings Singles Unidentified 2 1 1 4 3 1 2 4 2 2 3 1 5 2 2 1 1 1 3 1 4 Total 4 4 10 7 6 4 6 5 Grand Total 26 16 18 16 16 13 9 12 Snow was melting when dens were examined, and the ground at most den sites had several inches to 15 feet of snow. In most cases, bears tunneled through snow when emerging from dens. One atypical den formed entirely in snow by a single bear on the Alaska Peninsula had an S-shaped tunnel 19 feet long, a small alcove, and a cone-shaped chamber 9 feet high and 6 feet wide (Fig. 2A). Two dens, each occupied by a female with cubs, contained alder and willow branches. Den floors were evidently lined with branches during the final phase of construction in the fall. Three family groups and an unidentified bear had beds outside their dens. Some beds were in snow and some in soil. Figure 2C shows the orientation of beds used by a female with two yearlings. It appears bears constructed and used these beds after they first emerged and until they abandoned the den site. Other Denning Activities Investigators revisited 12 dens during succeeding fall and spring periods to determine if bears had used them again. There was only one instance of possible re-use. Most bears probably construct new dens each fall because thawing, seepage and erosion cause dens to collapse during the spring and summer. A few cases of denning in natural rock caves have been reported on Kodiak and the Alaska Peninsula, but none were found during this study. Denning could be expected in such rock caves during successive years. The one instance of co-occupancy, other than in dens used by females with young, was by two bears judged to be two-year-olds. These animals were probably litter mates that denned for the first time without their mother. No evidence of mortality associated with denning or parturition was discovered. 129
TABLE 3. DIMENSIONS IN FEET OF 26 BROWN BEAR DENS EXAMINED ON KODIAK ISLAND AND THE ALASKA PENINSULA. Excavated Material Entrance Tunnel Chamber Den Type Width Length Width Height Length Average Diameter Width Length and No. 1 4.8 Females w/cubs Mean A-1-65 A-5-67 A-1-68 A-2-68 A-3-68 A-5-68 2 6 8687 7 2 15 12 25 20 15 17 3.1 3.6 2.5 3.7 2.9 2.6 3.1 3.8 3.0 2.0 2.2 3.6 4.7 3.2 0 6.0 0 009.2 7.6 3 — — — — 2.3 3.2 2.7 5.0 ca. 3 4.8 4.3 5.0 5.0 4.5 8.0 ca. 4 5.4 11.9 9.2 5.0 7.2 Height ca. 4 3.3 3.9 3.8 4.3 4.0 Females w/yrlings Mean K-l-70 K-2-70 K-8-70 2 2 6 6 15 15 2 2 1.8 2.4 3.2 2.5 1.8 4.6 2.2 2.9 6.3 3.3 0 4.8 3.0 4.5 — 3.7 4.6 6.0 4.1 4.9 6.3 8.5 9.0 7.9 4.2 3.0 2.5 3.2 130
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IUCN Publications new series No. 23
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I n t he bear park on Hokkai do. (P
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Bear Proceedings © IUCN, Morges, S
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Contents Introduction to the biolog
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Introduction to the Biology and Man
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National parks in the United States
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Dr. Krott studied the behaviour of
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LENTFER, J. W., HENSEL, R. J., MILL
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PANEL 1: THE ECOLOGY, POPULATION CH
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dary. This procedure has the advant
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TABLE 2. ACTIVITY AREA DATA FROM FO
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found along adjacent foothills. Low
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numerous census techniques and in m
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SEX RATIOS There was no significant
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HARTLEY, H. O. 1958. Maximum likeli
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to the grizzlies, soapberry (Shephe
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In an area of approximately 500 squ
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cruises of known mileage by foot an
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156 miles on the Sugar lands route
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From a total of 92 survey forms dis
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Fig. 1 Location of standard canine
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TABLE 2. PRODUCT-MOMENT CORRELATION
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FIG. 2. Age structure of the 1967 b
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However, certain relationships were
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head from these bears. Commission p
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Fig. 1. Distribution of 1969 grizzl
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TABLE 4. ANNUAL HARVEST OF GRIZZLY
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on an offal pile or on a deer or el
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An undetermined species of hookworm
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Providing information to personnel
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Fig. 1 Distribution and abundance o
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still winter conditions. On emergin
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utting time were in an exhausted st
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ut recent information is lacking. F
Page 75 and 76: Simultaneously the isolated souther
Page 77 and 78: CONSERVATION The brown bear is not
Page 79 and 80: F. Craighead. In the Yellowstone po
Page 81 and 82: PANEL 2: DENNING—CONTROL MECHANIS
Page 83 and 84: Fig. 1 Radio location system Fig. 2
Page 85 and 86: neck diameter by inserting two scre
Page 87 and 88: Fig. 5 Bearings of transmitter sign
Page 89 and 90: TABLE 4. CHARACTERISTICS OF ACTIVE
Page 91 and 92: TABLE 5. CHARACTERISTICS OF INACTIV
Page 93 and 94: Fig. 7 Location of den of No. 40 by
Page 95 and 96: Fig. 10 Den of sow No. 40 (1966), s
Page 97 and 98: the cave, she could discourage unso
Page 99 and 100: emained in the den. In late afterno
Page 101 and 102: Our observations reveal that mature
Page 103 and 104: , , 1971. Biotelemetry Research wit
Page 105 and 106: winter denning of bears to allow an
Page 107 and 108: Fig. 1 View of den excavated by bla
Page 109 and 110: from the earth as a heat sink to th
Page 111 and 112: DENNING OF POLAR BEARS Conditions T
Page 113 and 114: Fig. 6 Polar bear maximum and minim
Page 115 and 116: and experiments of Whalen (1965) ar
Page 117 and 118: throughout the year on wild caught
Page 119 and 120: Fig. 9 EKG intervals of mammals. No
Page 121 and 122: SOUTH, F. E. & HOUSE, W. A. 1967. E
Page 123 and 124: willow and grasses in mountainous a
Page 125: TABLE 1. FREQUENCY DISTRIBUTION OF
Page 129 and 130: Fig. 2 Three den types: (A) atypica
Page 131 and 132: to 40°F; nice clear high pressure
Page 133 and 134: around that you could equate to a p
Page 135 and 136: PANEL 3: POLAR BEAR STUDIES Infra-r
Page 137 and 138: projection on a screen or on photog
Page 139 and 140: PANEL 3: POLAR BEAR STUDIES Further
Page 141 and 142: RESULTS A major polar bear winter d
Page 143 and 144: A small number of denned adults wit
Page 145 and 146: Fig. 6 A winter den of a female and
Page 147 and 148: ears has never been reported outsid
Page 149 and 150: gravel areas shallow dens are dug l
Page 151 and 152: Fig. 11 Diagram of a typical deep b
Page 153 and 154: DISCUSSION The winter polar bear de
Page 155 and 156: Even the winter dens differ from th
Page 157 and 158: Until 1970, the following major law
Page 159 and 160: He states that there is a connectio
Page 161 and 162: 7. LÖNÖ, O. 1970. Norsk Polarinst
Page 163 and 164: Polar pack ice covers the central p
Page 165 and 166: POLAR BEAR DISTRIBUTION IN RELATION
Page 167 and 168: from the vicinity of Wrangel Island
Page 169 and 170: PANEL 3: POLAR BEAR STUDIES Immobil
Page 171 and 172: After firing we kept watch over the
Page 173 and 174: RESULTS Results of the experiments
Page 175 and 176: } Duration of immobilization; comme
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One attempt was made to immobilize
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BREEDING ECOLOGY General informatio
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Fig. 1 The Drem-Head mountains, an
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TABLE 2. DISTRIBUTION OF DENS ACCOR
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Fig. 5 Structure of 6 dens: 1. (a)
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Fig. 7 Interior of den, showing cla
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We were lucky to observe three time
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In the upper reaches of the Neozhid
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In good weather, opened dens with s
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PANEL 3: POLAR BEAR STUDIES SUMMARY
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not be done, the hunter was given a
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Fig. 1 The contribution of year cla
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DISCUSSION Fluctuations in bear har
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REFERENCES FREE, S.& SAUER, P.R. 19
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the primate temporal lobe which is
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Fig. 1 The two cubs sucking on the
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Kit drank more on only 15. In June
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would immediately show interest, co
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Fig. 5 Resting self-play with branc
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A highly developed greeting ceremon
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KROTT, P. & KROTT, G. 1963. Zum Ver
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EVOLUTION OF BLACK BEARS AND THE BR
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they did not in general cease their
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Fig. 4 Fig. 3 226
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Many field observations suggest tha
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groups even with very frequent dump
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PANEL 4: BEAR BEHAVIOUR Social Beha
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The capture and tagging of 13 bears
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TABLE 3. SOCIAL RANK OF 22 BEARS OB
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TABLE 4. BEHAVIOR COMPONENTS OF BEA
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Fig 5 Bear 'confronting' an opponen
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Bears at the falls minimized compet
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the same manner as Mr. Stonorov obs
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males have their way. I'd say this,
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my way back to camp and I'd carried
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ears in Montana. It was masked by g
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ears like that, the subadults, usua
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The second observation I want to ma
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In the summer of 1969,Mr.Carlye Bla
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TABLE 1. DEMOGRAPHIC CHARACTERISTIC
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TABLE 3. KNOWLEDGE OF BLACK BEAR FO
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conception of the size of the black
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Occupation Occupation was not relat
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TABLE 6. ATTITUDE TOWARD ANIMALS IN
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The Elkmont campers were most conce
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TABLE 9. VISITOR ATTITUDE CONCERNIN
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TABLE 12. ANSWERS OF VISITORS TO QU
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PANEL 5: BEARS AND HUMAN BEINGS Pre
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number of visitors per year was abo
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TABLE 1. RELATIONSHIPS BETWEEN CAMP
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that fed at a dump. Weaned young, s
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TABLE 3. RECORDS ON GRIZZLY-CAUSED
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Canyon. Other coinciding circumstan
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number of intrusions into park deve
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REFERENCES CRAIGHEAD, J. J., HORNOC
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Glacier National Park Glacier Natio
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settlements in the park. Arthur Whe
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will be summarised here to provide
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C.P.R. 1889. Fishing and Shooting o
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Much later, in 1875, George Bird Gr
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its community. In other words, its
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COUES, Elliott (Ed.) 1965a. New lig
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PANEL 5: BEARS AND HUMAN BEINGS Gri
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to and after this month. Our census
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They have shaped and are integrally
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individually and quantitatively doc
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ecome thoroughly conditioned to man
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National Park Service Report, 'Griz
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This just equals the average annual
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has helped the sanitation problem.
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3 years of phase-out (Table 7). Thu
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TABLE 8. CAMPGROUND CAPTURES OF FIV
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TABLE 9. DISPERSAL OF 34 COLOR-MARK
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Fig. 6 Dispersal of 34 marked and t
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Fig. 10 A grizzly individually mark
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No. 8 moved 14 airline miles to the
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CRAIGHEAD, J. J. & CRAIGHEAD, F. C.
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PANEL 5: BEARS AND HUMAN BEINGS SUM
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ears present the least conflict wit
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een marked. This could be done alon
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spring and fall and so in a way it
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that we would have too much difficu
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and southward into the Mizo Hills a
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now but a relict population, is und
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This suggests a population of close
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Yoho, Kootenay and Glacier National
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support approximately 3, 300 bears
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Natural mortality is estimated at a
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TABLE 2. BLACK BEAR GAME STATUS STA
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TABLE 3. KILL FIGURES FOR BLACK BEA
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The research already undertaken has
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smallest takes is 11 bears from a t
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transplant or into zoos may have to
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HATLER,D. F. 1967. Some aspects in
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Index Figures in bold type are rele
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INDEX OF OTHERS WHO PARTICIPATED IN
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