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will be required before the technical practicality of the method under a wide range of conditions can be judged. The costs of buying or leasing imaging infra-red scanners and the expense of operating aircraft are substantial. Nevertheless, the initial testing indicates that this method is probably superior to any other and could have application in line transect sampling systems aimed at yielding estimates of bear numbers over fairly large regions. REFERENCES CROON, G. w., MCCULLOUGH, D. R., OLSON, C. E. Jr. & QUEAL, L. M. 1968. Infra-red Scanning Techniques for Big Game Censusing. J. Wildl. Mgmt. 32 (4): 751-759. THACKREY,D. E. 1968. Research in Infra-red Sensing. Research News. 18(8). Office of Research Administration, Univ. of Michigan, Ann Arbor. 12 pp. WOLFE, W. L. (Editor). 1965. Handbook of Military Infra-red Technology. U.S. Govt. Printing Office, Washington, D.C. 906 pp. 141
PANEL 3: POLAR BEAR STUDIES Further Notes on Polar Bear Denning Habits CHARLES J. JONKEL Canadian Wildlife Service, Ottawa GEORGE B. KOLENOSKY Ontario Department of Lands and Forests, Maple RICHARD J. ROBERTSON Manitoba Department of Lands and Forests, The Pas RICHARD H.RUSSELL Zoology Department, University of Alberta, Edmonton SUMMARY Polar bears construct maternity dens in the snow throughout their range. The Owl River maternity denning area on the Manitoba coast of Hudson Bay, Canada, had a measured productivity of 100-150 cubs in 1970 and 1971. Maternity denning is now confirmed for the Twin Islands in James Bay, but estimates of productivity for James Bay and the Ontario coast of Hudson Bay should still be made. Female polar bears build a variety of dens in the vicinity of their winter dens and along their route as they move to the sea ice. This makes the censusing of maternity dens and estimating of productivity difficult. The winter dens in Hudson and James bays differ from high arctic dens in that earth chambers are used, with snow dens added as winter progresses. Summer denning occurs along the Manitoba and Ontario coasts of Hudson Bay, and on the islands in James Bay. Surface pits, shallow dens and deep burrows are the three basic types of earth dens built. All three types appear to be constructed for temperature regulation by the bears, but each type is sometimes used later for shelter, protection from insects, protection from other bears or for winter dens. These behavioural adaptations appear significant in delineating a discrete polar bear population for James Bay and southern Hudson Bay. INTRODUCTION Polar bears (Ursus maritimus Phipps) den throughout their circum-polar range. Winter maternity dens constructed in drifted snow by pregnant females are the most common type and have been described by many arctic writers, especially Uspenski & Chernyavski (1965) & Harington (1968). Such dens are typically prepared by the female digging into a new drift, but sometimes she lies on or digs into a drift remnant from the previous year, allowing the snow to cover her. The maternity den is later expanded, perhaps by body heat, consumption of snow, scratching or digging. Maternity dens dug into deeply drifted areas show considerable evidence of design and construction (Uspenski & Chernyavski op. cit.; Harington op. cit.). Certain polar bears other than pregnant females construct winter dens in the snow, especially in maternity denning areas (Van de Velde 1957, 1971; Harington, op. cit.). Uspenski & Chernyavski (op. cit.) found no such evidence on Wrangel Island and the ecological and behavioural causes for this type of denning 142
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IUCN Publications new series No. 23
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I n t he bear park on Hokkai do. (P
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Bear Proceedings © IUCN, Morges, S
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Contents Introduction to the biolog
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Introduction to the Biology and Man
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National parks in the United States
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Dr. Krott studied the behaviour of
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LENTFER, J. W., HENSEL, R. J., MILL
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PANEL 1: THE ECOLOGY, POPULATION CH
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dary. This procedure has the advant
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TABLE 2. ACTIVITY AREA DATA FROM FO
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found along adjacent foothills. Low
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numerous census techniques and in m
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SEX RATIOS There was no significant
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HARTLEY, H. O. 1958. Maximum likeli
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to the grizzlies, soapberry (Shephe
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In an area of approximately 500 squ
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cruises of known mileage by foot an
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156 miles on the Sugar lands route
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From a total of 92 survey forms dis
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Fig. 1 Location of standard canine
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TABLE 2. PRODUCT-MOMENT CORRELATION
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FIG. 2. Age structure of the 1967 b
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However, certain relationships were
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head from these bears. Commission p
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Fig. 1. Distribution of 1969 grizzl
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TABLE 4. ANNUAL HARVEST OF GRIZZLY
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on an offal pile or on a deer or el
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An undetermined species of hookworm
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Providing information to personnel
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Fig. 1 Distribution and abundance o
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still winter conditions. On emergin
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utting time were in an exhausted st
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ut recent information is lacking. F
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Simultaneously the isolated souther
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CONSERVATION The brown bear is not
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F. Craighead. In the Yellowstone po
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PANEL 2: DENNING—CONTROL MECHANIS
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Fig. 1 Radio location system Fig. 2
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neck diameter by inserting two scre
Page 87 and 88: Fig. 5 Bearings of transmitter sign
Page 89 and 90: TABLE 4. CHARACTERISTICS OF ACTIVE
Page 91 and 92: TABLE 5. CHARACTERISTICS OF INACTIV
Page 93 and 94: Fig. 7 Location of den of No. 40 by
Page 95 and 96: Fig. 10 Den of sow No. 40 (1966), s
Page 97 and 98: the cave, she could discourage unso
Page 99 and 100: emained in the den. In late afterno
Page 101 and 102: Our observations reveal that mature
Page 103 and 104: , , 1971. Biotelemetry Research wit
Page 105 and 106: winter denning of bears to allow an
Page 107 and 108: Fig. 1 View of den excavated by bla
Page 109 and 110: from the earth as a heat sink to th
Page 111 and 112: DENNING OF POLAR BEARS Conditions T
Page 113 and 114: Fig. 6 Polar bear maximum and minim
Page 115 and 116: and experiments of Whalen (1965) ar
Page 117 and 118: throughout the year on wild caught
Page 119 and 120: Fig. 9 EKG intervals of mammals. No
Page 121 and 122: SOUTH, F. E. & HOUSE, W. A. 1967. E
Page 123 and 124: willow and grasses in mountainous a
Page 125 and 126: TABLE 1. FREQUENCY DISTRIBUTION OF
Page 127 and 128: TABLE 3. DIMENSIONS IN FEET OF 26 B
Page 129 and 130: Fig. 2 Three den types: (A) atypica
Page 131 and 132: to 40°F; nice clear high pressure
Page 133 and 134: around that you could equate to a p
Page 135 and 136: PANEL 3: POLAR BEAR STUDIES Infra-r
Page 137: projection on a screen or on photog
Page 141 and 142: RESULTS A major polar bear winter d
Page 143 and 144: A small number of denned adults wit
Page 145 and 146: Fig. 6 A winter den of a female and
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Page 149 and 150: gravel areas shallow dens are dug l
Page 151 and 152: Fig. 11 Diagram of a typical deep b
Page 153 and 154: DISCUSSION The winter polar bear de
Page 155 and 156: Even the winter dens differ from th
Page 157 and 158: Until 1970, the following major law
Page 159 and 160: He states that there is a connectio
Page 161 and 162: 7. LÖNÖ, O. 1970. Norsk Polarinst
Page 163 and 164: Polar pack ice covers the central p
Page 165 and 166: POLAR BEAR DISTRIBUTION IN RELATION
Page 167 and 168: from the vicinity of Wrangel Island
Page 169 and 170: PANEL 3: POLAR BEAR STUDIES Immobil
Page 171 and 172: After firing we kept watch over the
Page 173 and 174: RESULTS Results of the experiments
Page 175 and 176: } Duration of immobilization; comme
Page 177 and 178: One attempt was made to immobilize
Page 179 and 180: BREEDING ECOLOGY General informatio
Page 181 and 182: Fig. 1 The Drem-Head mountains, an
Page 183 and 184: TABLE 2. DISTRIBUTION OF DENS ACCOR
Page 185 and 186: Fig. 5 Structure of 6 dens: 1. (a)
Page 187 and 188: Fig. 7 Interior of den, showing cla
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We were lucky to observe three time
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In the upper reaches of the Neozhid
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In good weather, opened dens with s
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PANEL 3: POLAR BEAR STUDIES SUMMARY
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not be done, the hunter was given a
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Fig. 1 The contribution of year cla
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DISCUSSION Fluctuations in bear har
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REFERENCES FREE, S.& SAUER, P.R. 19
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the primate temporal lobe which is
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Fig. 1 The two cubs sucking on the
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Kit drank more on only 15. In June
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would immediately show interest, co
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Fig. 5 Resting self-play with branc
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A highly developed greeting ceremon
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KROTT, P. & KROTT, G. 1963. Zum Ver
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EVOLUTION OF BLACK BEARS AND THE BR
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they did not in general cease their
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Fig. 4 Fig. 3 226
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Many field observations suggest tha
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groups even with very frequent dump
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PANEL 4: BEAR BEHAVIOUR Social Beha
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The capture and tagging of 13 bears
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TABLE 3. SOCIAL RANK OF 22 BEARS OB
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TABLE 4. BEHAVIOR COMPONENTS OF BEA
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Fig 5 Bear 'confronting' an opponen
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Bears at the falls minimized compet
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the same manner as Mr. Stonorov obs
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males have their way. I'd say this,
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my way back to camp and I'd carried
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ears in Montana. It was masked by g
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ears like that, the subadults, usua
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The second observation I want to ma
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In the summer of 1969,Mr.Carlye Bla
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TABLE 1. DEMOGRAPHIC CHARACTERISTIC
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TABLE 3. KNOWLEDGE OF BLACK BEAR FO
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conception of the size of the black
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Occupation Occupation was not relat
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TABLE 6. ATTITUDE TOWARD ANIMALS IN
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The Elkmont campers were most conce
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TABLE 9. VISITOR ATTITUDE CONCERNIN
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TABLE 12. ANSWERS OF VISITORS TO QU
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PANEL 5: BEARS AND HUMAN BEINGS Pre
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number of visitors per year was abo
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TABLE 1. RELATIONSHIPS BETWEEN CAMP
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that fed at a dump. Weaned young, s
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TABLE 3. RECORDS ON GRIZZLY-CAUSED
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Canyon. Other coinciding circumstan
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number of intrusions into park deve
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REFERENCES CRAIGHEAD, J. J., HORNOC
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Glacier National Park Glacier Natio
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settlements in the park. Arthur Whe
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will be summarised here to provide
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C.P.R. 1889. Fishing and Shooting o
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Much later, in 1875, George Bird Gr
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its community. In other words, its
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COUES, Elliott (Ed.) 1965a. New lig
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PANEL 5: BEARS AND HUMAN BEINGS Gri
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to and after this month. Our census
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They have shaped and are integrally
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individually and quantitatively doc
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ecome thoroughly conditioned to man
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National Park Service Report, 'Griz
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This just equals the average annual
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has helped the sanitation problem.
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3 years of phase-out (Table 7). Thu
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TABLE 8. CAMPGROUND CAPTURES OF FIV
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TABLE 9. DISPERSAL OF 34 COLOR-MARK
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Fig. 6 Dispersal of 34 marked and t
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Fig. 10 A grizzly individually mark
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No. 8 moved 14 airline miles to the
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CRAIGHEAD, J. J. & CRAIGHEAD, F. C.
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PANEL 5: BEARS AND HUMAN BEINGS SUM
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ears present the least conflict wit
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een marked. This could be done alon
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spring and fall and so in a way it
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that we would have too much difficu
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and southward into the Mizo Hills a
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now but a relict population, is und
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This suggests a population of close
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Yoho, Kootenay and Glacier National
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support approximately 3, 300 bears
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Natural mortality is estimated at a
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TABLE 2. BLACK BEAR GAME STATUS STA
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TABLE 3. KILL FIGURES FOR BLACK BEA
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The research already undertaken has
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smallest takes is 11 bears from a t
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transplant or into zoos may have to
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HATLER,D. F. 1967. Some aspects in
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Index Figures in bold type are rele
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INDEX OF OTHERS WHO PARTICIPATED IN
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