Ohio subset of Plant Communities of the Midwest ... - NatureServe
Ohio subset of Plant Communities of the Midwest ... - NatureServe
Ohio subset of Plant Communities of the Midwest ... - NatureServe
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Open and Emergent Marshes: Eastern Emergent Marshes<br />
Typha spp. - Schoenoplectus acutus - Mixed Herbs <strong>Midwest</strong> Herbaceous Vegetation<br />
Cattail Species - Hardstem Bulrush - Mixed Herbs <strong>Midwest</strong> Herbaceous Vegetation<br />
<strong>Midwest</strong> Mixed Emergent Deep Marsh<br />
CEGL002229<br />
DESCRIPTION: This deepwater emergent marsh community is dominated by perennial herbaceous vegetation with<br />
graminoid leaves. A typical example <strong>of</strong> this marsh contains a mosaic <strong>of</strong> emergents, submergents, and floating-leaved<br />
plants interspersed with areas <strong>of</strong> open water (Harris et al. 1996). Various kinds <strong>of</strong> emergents may dominate a marsh<br />
depending on <strong>the</strong> water depth. Quite <strong>of</strong>ten <strong>the</strong> vegetation arranges itself in belts (wetland zonation), with a particular<br />
species or range <strong>of</strong> species occupying specific depths from <strong>the</strong> shoreline to deep open water. Marshes may display<br />
areas <strong>of</strong> open water, but vegetation dominates (>30% cover). Typha latifolia, Typha angustifolia, and Schoenoplectus<br />
acutus (= Scirpus acutus) dominate this dynamic ecosystem. Sedges are also common (e.g. Carex lupuliformis and<br />
Carex hyalinolepis in <strong>the</strong> eastern part <strong>of</strong> <strong>the</strong> range). A diverse assemblage <strong>of</strong> grasses, floating-leaved aquatics, and<br />
submerged aquatics are present (TNC 1995a).<br />
These highly productive wetlands are found in glacial potholes, river valleys, ponds, and on lakeplains. They are<br />
characterized by continuous inundation and are considered a deep marsh. Water depth averages 0.3-0.6 m, ranging<br />
from several centimeters to more than one meter for a significant part <strong>of</strong> <strong>the</strong> growing season. Seasonal flooding during<br />
winter and spring or flooding during heavy rains help maintain <strong>the</strong>se marshes by causing water exchange which<br />
replenishes freshwater and circulates nutrients and organic debris. Soils can be mineral or organic but are saturated,<br />
flooded, or ponded long enough during <strong>the</strong> growing season to develop anaerobic conditions in <strong>the</strong> upper part.<br />
Vegetative diversity and density are highly variable in response to water depth, water chemistry, and natural forces.<br />
Seasonal flooding and heavy rains influence vegetative growth, aquatic animals, and nutrient cycling in marsh ecosystems.<br />
COMMENTS: 2, MCS. This community type is very broadly defined. It can be highly dynamic from one season to <strong>the</strong><br />
next, and species composition and density are very dependent on water regime and sometimes <strong>the</strong> mineral content <strong>of</strong><br />
water and soil (Niering 1985). Deep marsh is defined as having water depths <strong>of</strong> 15 cm to over 1 m throughout all or most<br />
<strong>of</strong> <strong>the</strong> growing season (Shaw and Fredine 1971, in Eggers and Reed 1987). The concept and distribution <strong>of</strong> this<br />
community in <strong>the</strong> Sou<strong>the</strong>ast needs reassessment; it should be compared to Typha latifolia Sou<strong>the</strong>rn Herbaceous<br />
Vegetation (CEGL004150), particularly in <strong>the</strong> Mississippi alluvial plain <strong>of</strong> Missouri (M. Leahy pers. comm. 1999). Many <strong>of</strong><br />
<strong>the</strong> presettlement occurrences <strong>of</strong> this community have been drained and converted to cropland or destroyed by siltation,<br />
which greatly accelerates <strong>the</strong> natural successional process from shallow inundation to moist soil. Lythrum salicaria is an<br />
aggressive exotic species that threatens this community in Canada, <strong>the</strong> Nor<strong>the</strong>ast, and more recently in <strong>the</strong> <strong>Midwest</strong>.<br />
Stands in Province 212 may deserve to be recognized as a separate nor<strong>the</strong>rn type, but fur<strong>the</strong>r data are needed.<br />
CONSERVATION RANK: G4. This type is subject to many forms <strong>of</strong> human impact, particularly draining for farmland.<br />
DISTRIBUTION: This community is found in glacial potholes, river valleys, ponds, and on lake plains throughout <strong>the</strong><br />
midwestern and parts <strong>of</strong> <strong>the</strong> eastern United States, ranging from <strong>Ohio</strong> and Ontario west to North Dakota, south to<br />
Nebraska and east to Alabama.<br />
USFS ECOREGIONS: 212He:CCC, 212Hj:CCP, 212Ho:CCC, 212Hq:CCP, 212Ht:CCC, 212Hv:CCC, 212Ib:C, 212Ja:CPP,<br />
212Jb:CP, 212Jc:CP, 212Jf:CPP, 212Jj:CPP, 212Jl:CPP, 212Jm:CPP, 212Jn:CP, 212Jo:CP, 212Ka:CPP, 212Mb:CPP,<br />
212Na:CCP, 212Nb:CCP, 212Nc:CCC, 212Nd:CC, 222A:CP, 222Cg:CCP, 222D:CP, 222E:CP, 222G:CP, 222Ha:CCC,<br />
222Hb:CCC, 222Ia:CCC, 222Jc:CCC, 222Jg:CCC, 222Jh:CCC, 222Ji:CCC, 222Jj:CCC, 222Kd:CCC, 222Ke:CCC, 222Kf:CCC,<br />
222Kg:CCC, 222Kj:CCC, 222Mc:CCC, 222Md:CCC, 222Me:CCC, 222Na:CCC, 234A:PP, 251Aa:CCC, 251Ab:CCC,<br />
251Bb:CCC, 251Bd:CCC, 251Be:CCC, 251Cb:CCC, 251Cf:CCC, 251Cg:CCC, 251Ch:CCC, 251Cm:CCC, 251Cp:CCC,<br />
251Dg:CCC, 251Eb:CCC<br />
CONSERVATION REGIONS: 35:C, 36:C, 37:P, 43:C, 44:P, 46:C, 47:C, 48:C, 50:<br />
STATES: AL IA IL IN KY MI MN MO ND NE OH SD TN WI PROVINCES: ON<br />
MIDWEST HERITAGE SYNONYMY: IL marsh (S); marsh (N) I<br />
IN marsh +<br />
MI inland emergent marsh +<br />
MN mixed emergent marsh (prairie section); mixed emergent marsh (forest section) I<br />
MO freshwater marsh +<br />
NE freshwater marsh +<br />
OH mixed emergent marsh; mixed emergent riverine community I<br />
WI deep marsh (mixed emergent subtype) =<br />
OTHER SYNONYMY: Marsh: Mixed: Mineral Substrate (W5) (Harris et al. 1996) B, Marsh: Mixed: Organic Substrate<br />
(W10) (Harris et al. 1996) B<br />
USNVC HIERARCHY: TYPHA (ANGUSTIFOLIA, LATIFOLIA) - (SCHOENOPLECTUS SPP.) SEMIPERMANENTLY FLOODED HERBACEOUS<br />
ALLIANCE (V.A.5.N.l)<br />
PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS<br />
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