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Ohio subset of Plant Communities of the Midwest ... - NatureServe

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Wooded Swamps and Floodplains: <strong>Midwest</strong>ern Wet Flatwoods<br />

Fagus grandifolia - Acer saccharum - Quercus bicolor - Acer rubrum Flatwoods Forest<br />

American Beech - Sugar Maple - Swamp White Oak - Red Maple Flatwoods Forest<br />

Beech - Hardwoods Till Plain Flatwoods<br />

CEGL005173<br />

DESCRIPTION: This community is dominated by trees that can be found in both upland and lowland sites. Among <strong>the</strong><br />

most common are Acer rubrum, Acer saccharum, Fagus grandifolia, Quercus bicolor, Quercus macrocarpa, and<br />

Ulmus rubra. Associates include Carya ovata, Carya cordiformis, Celtis occidentalis, Fraxinus pennsylvanica,<br />

Liriodendron tulipifera, Quercus rubra, Quercus shumardii, and Ulmus americana. Typical shrub species include<br />

Asimina triloba, Euonymus obovata, and Lindera benzoin. The herbaceous layer includes Arisaema triphyllum (=<br />

Arisaema atrorubens), Actaea pachypoda, Circaea X intermedia (= Circaea canadensis), Galium spp., and Pilea<br />

pumila, among o<strong>the</strong>rs (Homoya et al. 1988, Anderson 1996).<br />

This community is found on flat to gently sloping sites on till plains. Soils are imperfectly to poorly drained and<br />

aerated. Soil types include Crosby, Blount, Fincastle, Brookston, and Pewamo (Homoya et al 1988)<br />

COMMENTS: 2, MCS. The type is based on <strong>the</strong> concept <strong>of</strong> <strong>the</strong> Indiana state type - central till plain flatwoods (Homoya<br />

et al. 1988). This type occurs as a mosaic <strong>of</strong> upland rises with Fagus grandifolia and o<strong>the</strong>r hardwoods and wetland<br />

areas with Quercus palustris, Quercus bicolor, Acer rubrum, and o<strong>the</strong>r wetland hardwoods. It is possible that <strong>the</strong><br />

mosaic may become extensive enough in some areas that <strong>the</strong> two parts could be treated separately, but this needs<br />

field checking. In any case, no o<strong>the</strong>r states or provinces report this kind <strong>of</strong> patterning. To help define <strong>the</strong> type, stands<br />

may require at least 25% beech basal area or cover to fit this type (Anderson 1996). In Ontario <strong>the</strong> type may occur<br />

above <strong>the</strong> escarpment in <strong>the</strong> Niagara Peninsula, but <strong>the</strong>se may have been treated as part <strong>of</strong> <strong>the</strong> Great Lakes<br />

lakeplain type, Quercus palustris - Quercus bicolor - Acer rubrum Flatwoods Forest (CEGL005037).<br />

CONSERVATION RANK: G2G3. There are probably fewer than 25 occurrences <strong>of</strong> this community rangewide; it is only<br />

known from central Indiana, where it is ranked S2, and from sou<strong>the</strong>astern Michigan and nor<strong>the</strong>astern <strong>Ohio</strong>. Currently<br />

19 occurrences have been documented from Indiana. There are probably fewer than 1000 acres <strong>of</strong> this community<br />

rangewide. Currently over 175 acres have been documented (some sites lack acreage data); <strong>the</strong> average size<br />

documented is 35 acres. Historical acreage is unknown. Many sites have been degraded by grazing and selective<br />

logging, and <strong>the</strong>re has probably been some overall reduction in acreage. This community has somewhat restricted<br />

environmental requirements: it is found over till plains with imperfectly to poorly drained and aerated soils. The range<br />

is somewhat restricted; it is reported from four subsections in one ecoregion province.<br />

DISTRIBUTION: This community is found in <strong>the</strong> central midwestern United States and adjacent Canada, ranging from<br />

central Indiana, western <strong>Ohio</strong>, and sou<strong>the</strong>astern Michigan to possibly adjacent Ontario.<br />

USFS ECOREGIONS: 222Ha:CCC, 222Hb:CCC, 222Hf:CCC, 222If:CCC<br />

CONSERVATION REGIONS: 45:C<br />

STATES: IN MI OH PROVINCES: ON<br />

MIDWEST HERITAGE SYNONYMY: IN central till plain flatwoods =<br />

MI pin oak depression<br />

OH beech-oak-red maple forest <br />

OTHER SYNONYMY: South LaPorte Woods (Lindsey et al. 1969) =, Wet Beech Forests (Anderson 1996) B. Anderson<br />

describes <strong>the</strong> differences between Wet Beech Forest stands in southwestern <strong>Ohio</strong> from those in <strong>the</strong> west/northwest<br />

part <strong>of</strong> <strong>Ohio</strong>. Some <strong>of</strong> Anderson's mixed swamp type may also fall here, depending on percentage <strong>of</strong> beech used to<br />

separate his wet beech forest type from this type.<br />

USNVC HIERARCHY: FAGUS GRANDIFOLIA - QUERCUS SPP. - ACER SPP. FOREST ALLIANCE (I.B.2.N.a)<br />

PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS<br />

187

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