Forests and Woodlands: <strong>Midwest</strong>ern Dry and Dry-mesic Oak Forests Quercus velutina - Quercus alba - Carya (glabra, ovata) Forest Black Oak - White Oak - (Pignut Hickory, Shagbark Hickory) Forest Black Oak - White Oak - Hickory Forest CEGL002076 DESCRIPTION: Trees in this community <strong>of</strong>ten have moderate to short trunks and spreading crowns, and canopy can vary from open to closed (50-100%). Quercus velutina, Quercus alba, Carya glabra, and Carya ovata are typical tree dominants. Associated oaks can include Quercus ellipsoidalis (northward), Quercus muehlenbergii, and southward, Quercus coccinea, Quercus prinus and Quercus stellata. Typical shrubs and small trees include Cornus florida (southward), Cornus foemina, Corylus americana (northward), Ostrya virginiana, and Sassafras albidum. Vines include Toxicodendron radicans, Par<strong>the</strong>nocissus quinquefolia, and Vitis spp. The herbaceous layer can include Agrimonia gryposepala, Agrimonia rostellata, Amphicarpaea bracteata, Botrychium virginianum, Carex blanda, Carex pensylvanica, Desmodium glutinosum, Desmodium nudiflorum, Dioscorea quaternata, Galium circaezans, Geranium maculatum, Polystichum acrostichoides, Maian<strong>the</strong>mum racemosum (= Smilacina racemosa), and Maian<strong>the</strong>mum stellatum (= Smilacina stellata), among o<strong>the</strong>rs (Curtis 1959, White and Madany 1978, Nelson 1985, TNC 1995a). This community is <strong>of</strong>ten found on ridge crests or upper slopes that are well-drained to excessively drained. Soils are <strong>of</strong>ten sandy loam, thin and rocky, with outcroppings <strong>of</strong> exposed bedrock. Bedrock is sandstone, siltstone, chert, or shale, or northward covered by thin loess or glacial till (Curtis 1959, White and Madany 1978, Nelson 1985). Occasional drought stress, wind, and lightning damage are evident. Historically, fire may have periodically affected this community, increasing its range into more moist sites (Fralish et al. 1991, Robertson and Heikens 1994). Fires in <strong>the</strong> Shawnee and Ozark Hills, just north <strong>of</strong> <strong>the</strong> range <strong>of</strong> this type, were more frequent (almost annual) from <strong>the</strong> early 1900s to 1930, but <strong>the</strong>re is little information on presettlement (prior to 1800) fire frequency (Robertson and Heikens 1994). Erosion also converts mesic forest soils to dry-mesic, <strong>the</strong>reby creating conditions which favor <strong>the</strong> occurrence <strong>of</strong> this community. McCune and Cottam (1985) present a detailed 30-year successional analysis <strong>of</strong> a stand in sou<strong>the</strong>rn Wisconsin. COMMENTS: 2, MCS. It is possible that unglaciated stands may differ sufficiently from glaciated stands to warrant separating into two types, but this depends on resolving <strong>the</strong> types distribution southward. Is <strong>the</strong>re a thin acid soil, non-glaciated type (i.e., Quercus alba - Carya alba - (Quercus velutina) / Polystichum acrostichoides - (Carex picta) Forest (CEGL007795)) versus deeper, glacial soil type This type appears to go as far north as nor<strong>the</strong>rn Illinois, Indiana, and sou<strong>the</strong>rn Michigan, and probably into sou<strong>the</strong>rn Wisconsin in section 222K, but not 222L (H. Dunevitz pers. comm. 2000, Eric Epstein pers. comm. 1999). The type concept in Wisconsin remains to be resolved. Indiana suggested that <strong>the</strong> unglaciated stands have more Quercus coccinea. In Michigan this could fit <strong>the</strong> interlobate region. Types on sand and typically more dominated by Quercus velutina are placed in ei<strong>the</strong>r Quercus velutina - Quercus alba / Vaccinium (angustifolium, pallidum) / Carex pensylvanica Forest (CEGL005030) or Quercus velutina / Carex pensylvanica Forest (CEGL002078). This type has been described in sou<strong>the</strong>rn Illinois by Robertson et al. (1984), Fralish (1988), and Fralish et al. (1991). Braun (1950, p. 145-146) also noted <strong>the</strong> prominence <strong>of</strong> black and white oaks in <strong>the</strong> Ozark Hills and Illinois, <strong>the</strong> Mammoth Cave area <strong>of</strong> Kentucky, and throughout <strong>the</strong> oak - hickory forest region, especially <strong>the</strong> Mississippi Valley and Prairie Peninsula regions. In Minnesota, <strong>the</strong> Bigwoods Sou<strong>the</strong>ast section may not contain consistent enough black oak to fit this type, but nor<strong>the</strong>rn pin oak may be common. In <strong>the</strong> Ozarks this type may exist, but, based on <strong>the</strong> recommendations from D. Ladd, T. Nigh, D. Zollner, and B. Heumann, stands are placed ei<strong>the</strong>r in Quercus alba - Quercus stellata - Quercus velutina / Schizachyrium scoparium Woodland (CEGL002150) or in Quercus velutina - Quercus coccinea - Carya texana Ozark Forest (CEGL002399). Quercus alba - (Quercus velutina) - Carya ovata / Ostrya virginiana Forest (CEGL002011) may be equivalent to this type, at least in nor<strong>the</strong>rn Missouri (M. Leahy pers. comm. 1999) CONSERVATION RANK: G4. DISTRIBUTION: This oak forest community is found throughout <strong>the</strong> nor<strong>the</strong>rn and central midwestern United States and adjacent Canada, ranging from <strong>Ohio</strong> and Ontario, west to possibly sou<strong>the</strong>rn Wisconsin, south to nor<strong>the</strong>rn Missouri, and east to Indiana and possibly Kentucky. USFS ECOREGIONS: 221:P, 222Ca:CCC, 222De:CCC, 222Df:CCC, 222Dh:CCP, 222Di:CCP, 222Ek:CCC, 222Em:CCC, 222Fe:CCC, 222Ga:CCC, 222Gb:CCC, 222Gd:CCC, 222Ha:CCC, 222Hb:CCP, 222Hf:CCP, 222Jb:CCP, 222Jg:CCC, 222Jh:CCC, 222Ji:CCP, 222Jj:CCC, 222Kf:CCC, 222Kj:CCC, 231:C, 251Cf:CCC, 251Ci:CCC, 251Cj:CCC, 251Ck:CCC, 251Dc:CCC, 251Dd:CCC, 251Df:CCC, 251Dh:CCP CONSERVATION REGIONS: 36:C, 44:P, 46:C, 48:C STATES: AR IA IL IN KY MI MO OH PROVINCES: ON PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS 412
MIDWEST HERITAGE SYNONYMY: IL dry-mesic upland forest (S); dry upland forest (N) I IN dry upland forest + MI dry sou<strong>the</strong>rn forest - black oak-white oak + MO dry forest; dry acid bedrock forest (dry chert forest, dry sandstone forest, dry igneous forest) I OH oak-hickory forest OTHER SYNONYMY: Dry-mesic Upland Forest B. (IL) The Illinois Natural Areas Inventory names communities according to broad environmental conditions, Quercus velutina - Q. alba upper slope community type (Robertson et al. 1984) =. Good fit, White Oak-Black Oak-Hickory Association (Gordon 1969) =. Uncertain if equivalent. In <strong>Ohio</strong>, Gordon (1969, p. 40) notes that "Over <strong>the</strong> glacial till plains <strong>of</strong> <strong>Ohio</strong>, in <strong>the</strong> vicinity <strong>of</strong> prairie grasslands, forests were mainly <strong>of</strong> <strong>the</strong> white oak-black oak-shagbark hickory type at <strong>the</strong> time <strong>of</strong> settlement.", Oak-hickory forests (Anderson 1996) B. Anderson notes that white oak-red oak dominance may occur, and that his Oak-Hickory Forests type includes Gordon's white oak-black oak-shagbark hickory type., Sou<strong>the</strong>rn Dry Forest (Curtis 1959) =, TIB4aII4c. Quercus alba - Quercus velutina - Quercus falcata (Foti et al. 1994), Dry Forest, Dry Acid (Chert, Igneous, Sandstone) Forest (Nelson 1987) USNVC HIERARCHY: QUERCUS VELUTINA - QUERCUS ALBA - (QUERCUS COCCINEA) FOREST ALLIANCE (I.B.2.N.a) PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS 413
- Page 1 and 2:
PLANT COMMUNITIES OF THE MIDWEST CL
- Page 3 and 4:
CONTENTS APPENDIX: PLANT COMMUNITY
- Page 5 and 6:
OVERVIEW APPENDIX: PLANT COMMUNITY
- Page 7 and 8:
− The USFS (U.S. Forest Service)
- Page 9 and 10:
Box 2. State Natural Heritage progr
- Page 11 and 12:
INDEX TO ASSOCIATIONS BY ECOLOGICAL
- Page 13:
2.3.4.4. Interior Highlands Shale G
- Page 16 and 17:
Acid Peatlands: Northern Shrub/Gram
- Page 18 and 19:
Rich Peat Fens: Northeastern Rich F
- Page 20 and 21:
Rich Peat Fens: Midwestern Prairie
- Page 22 and 23:
Seeps: Midwestern Seepage Meadows S
- Page 24 and 25:
Open and Emergent Marshes: Eastern
- Page 26 and 27:
Open and Emergent Marshes: Eastern
- Page 28 and 29:
Open and Emergent Marshes: Eastern
- Page 30 and 31:
* Nonstandard type (needs review) G
- Page 32 and 33:
Open and Emergent Marshes: Great La
- Page 34 and 35:
Wet Prairies and Wet Meadows: Great
- Page 36 and 37:
Wet Prairies and Wet Meadows: Midwe
- Page 38 and 39: Wet Prairies and Wet Meadows: Midwe
- Page 40 and 41: Wooded Swamps and Floodplains: Nort
- Page 42 and 43: Wooded Swamps and Floodplains: Nort
- Page 44 and 45: Wooded Swamps and Floodplains: Nort
- Page 46 and 47: Wooded Swamps and Floodplains: Midw
- Page 48 and 49: Wooded Swamps and Floodplains: Midw
- Page 50 and 51: Wooded Swamps and Floodplains: Midw
- Page 52 and 53: Wooded Swamps and Floodplains: Midw
- Page 54 and 55: Wooded Swamps and Floodplains: Midw
- Page 56 and 57: Wooded Swamps and Floodplains: Midw
- Page 58 and 59: Wooded Swamps and Floodplains: Midw
- Page 60 and 61: Wooded Swamps and Floodplains: Appa
- Page 62 and 63: Wooded Swamps and Floodplains: Sout
- Page 64 and 65: Wooded Swamps and Floodplains: Sout
- Page 66 and 67: Shoreline Sand/Mud Strands, Beaches
- Page 68 and 69: Sphagnum wulfianum, Sphagnum warnst
- Page 70 and 71: Shoreline Sand/Mud Strands, Beaches
- Page 72 and 73: Rocky Shores: Great Lakes Alkaline
- Page 74 and 75: Rocky Shores: Appalachian Highlands
- Page 76 and 77: Rocky Uplands (Glades, Rock Barrens
- Page 78 and 79: DISTRIBUTION: This limestone glade
- Page 80 and 81: DISTRIBUTION: This graminoid shale
- Page 82 and 83: Cliffs, Talus, Buttes and Badlands:
- Page 84 and 85: Cliffs, Talus, Buttes and Badlands:
- Page 86 and 87: Cliffs, Talus, Buttes and Badlands:
- Page 90 and 91: Forests and Woodlands: Midwestern D
- Page 92 and 93: Forests and Woodlands: Midwestern M
- Page 94 and 95: Forests and Woodlands: Appalachian
- Page 96 and 97: Forests and Woodlands: Appalachian
- Page 98 and 99: Forests and Woodlands: Appalachian
- Page 100 and 101: Forests and Woodlands: Appalachian
- Page 102 and 103: Forests and Woodlands: Interior Hig
- Page 104 and 105: Savannas and Non-Rock Barrens: Midw
- Page 106 and 107: Savannas and Non-Rock Barrens: Midw
- Page 108 and 109: Prairies/Grasslands: Midwestern San
- Page 110 and 111: Prairies/Grasslands: Midwestern Thi
- Page 112 and 113: Prairies/Grasslands: Midwestern Dee