DISTRIBUTION: This limestone glade or barrens community is found in <strong>the</strong> central and eastern United States, ranging from sou<strong>the</strong>rn Illinois, Kentucky, Tennessee and Alabama, east to Georgia, western Virginia, West Virginia, and <strong>Ohio</strong>. USFS ECOREGIONS: 222Aq:CCC, 222De:CCC, 222Df:CCC, 222Dh:CCC, 222Di:CCC, 222Ei:CCC, 222Ek:CCC, 222El:CCC, 222Fc:CCC, 222Fd:CCC, 222Fe:CCC, 251Cf:CCC, 251Ci:CCC CONSERVATION REGIONS: 36:C, 38:C, 43:, 44:C, 49:, 50:C, 59: STATES: AL GA IL IN KY OH TN VA WV PROVINCES: MIDWEST HERITAGE SYNONYMY: IL limestone glade = IN limestone glade = OH little bluestem prairie + OTHER SYNONYMY: Xeric Limestone Prairie (Baskin et al. 1994), Barrens type (Hutchison et al. 1986). is sometimes treated broadly as a barrens type, Barrens type (Hutchison 1994). is sometimes treated broadly as a barrens type USNVC HIERARCHY: (JUNIPERUS VIRGINIANA) / SCHIZACHYRIUM SCOPARIUM - (BOUTELOUA CURTIPENDULA) WOODED HERBACEOUS ALLIANCE (V.A.6.N.q) PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS 304
Rocky Uplands (Glades, Rock Barrens, Outcrops and Alvars): Interior Highlands Shale Glades and Barrens Quercus marilandica - (Juniperus virginiana) / Schizachyrium scoparium - Danthonia spicata Wooded Herbaceous Vegetation Blackjack Oak - (Eastern Red-cedar) / Little Bluestem - Poverty Oatgrass Wooded Herbaceous Vegetation Central Shale Glade CEGL002428 DESCRIPTION: Trees can be found scattered or in isolated clumps and patches throughout <strong>the</strong> occurrence, along with a few saplings and shrubs. Quercus marilandica and Quercus stellata are <strong>the</strong> dominant trees (Quercus prinus is dominant in Indiana examples), and Pinus virginiana or o<strong>the</strong>r Quercus spp. may also be found in some examples. Vaccinium arboreum, Rhus aromatica and Ulmus alata are common shrubs. Vaccinium pallidum and Gaylussacia baccata are common in Indiana examples. <strong>Ohio</strong> stands also contain Cercis canadensis, Amelanchier arborea, Frangula caroliniana (= Rhamnus caroliniana), Rosa carolina, and Vaccinium stamineum. Schizachyrium scoparium and Danthonia spicata dominate <strong>the</strong> herbaceous layer, along with a diverse assemblage <strong>of</strong> forbs. Missouri examples, where <strong>the</strong> shales are fairly 'limey,' contain Astragalus distortus, Dalea purpurea, Porteranthus stipulatus, and Psoralidium tenuiflorum. The herbaceous flora <strong>of</strong> Indiana Interior Low Plateau examples (Siltstone Glades) includes Agalinis sp., Symphyotrichum patens (= Aster patens), Symphyotrichum shortii (= Aster shortii), Brickellia eupatorioides (= Kuhnia eupatorioides), Cunila origanoides, Dichan<strong>the</strong>lium depauperatum, Euphorbia corollata, Helianthus sp., Hypericum crux-andreae, Hypericum hypericoides, Liatris squarrosa, Manfreda virginica, Porteranthus stipulatus, Schizachyrium scoparium, and Tephrosia virginiana. Some woody plants recorded at <strong>the</strong>se sites include Pinus virginiana, Quercus prinus, Quercus stellata, Quercus marilandica, and Juniperus virginiana. In <strong>Ohio</strong>, Carex hirsutella, Agrostis perennans, Allium cernuum, Sericocarpus asteroides (= Aster paternus), Agalinis tenuifolia (= Gerardia tenuifolia), Houstonia canadensis (= Hedyotis canadensis), and Lithospermum canescens, among o<strong>the</strong>rs, may occur. Some grasses and forbs from <strong>the</strong> Ouachita Mountain examples include Agrostis elliottiana, Oeno<strong>the</strong>ra linifolia, Astragalus distortus, and Ruellia humilis. Lichens and mosses occur in pr<strong>of</strong>usion on exposed bedrock surfaces and on soils not covered with organic debris (leaf litter, wood) (White and Madany 1978, Nelson 1985, Homoya 1994, TNC 1995a, D. Minney pers. comm. 2000). In Missouri, this community occurs on moderate to steep slopes <strong>of</strong> dissected drainages along major streams and mounds, and shales have a strong limestone component, making <strong>the</strong>m less acid. In several square km <strong>of</strong> Illinois, a series <strong>of</strong> steep ridges occurs in thick shale outcrops. Shale glades occur <strong>the</strong>re on exposed slopes. The soil <strong>of</strong> <strong>the</strong>se communities is absent to thin (0-40 cm) and somewhat rapidly to very rapidly drained. The parent material is shale, with siltstone present in Indiana occurrences. Shale fragments and exposed bedrock are common on <strong>the</strong> surface. In <strong>Ohio</strong>, sites occur sparingly on estill shales with extreme southwestern exposures (White and Madany 1978, Nelson 1985, Homoya 1994, D. Minney pers. comm. 2000). Rapid wea<strong>the</strong>ring and erosion on steep slopes appear to be necessary for shale glades to persist. Fires may have been very occasional (Nelson 1985). COMMENTS: 2, MCS. This community has a restricted distribution and occurrences are small. In Indiana, Quercus prinus is <strong>the</strong> major dominant. Similar communities occur in West Virginia, and also with Quercus prinus as <strong>the</strong> primary scattered tree. In <strong>the</strong> Knobs region <strong>of</strong> Kentucky this community is threatened by strip mining. Total vegetative cover may sometimes be less than 10%. Shale glades in Missouri are so lime-rich that <strong>the</strong>y may be treated as a variant <strong>of</strong> limestone glade (M. Leahy pers. comm. 1999). In <strong>Ohio</strong>, areas <strong>of</strong> shale that have been plowed have come back as "cedar barrens" but are typically dominated by a variety <strong>of</strong> annuals (though most <strong>of</strong> <strong>the</strong>se are present at some level in <strong>the</strong> natural shale glades as well), and <strong>the</strong>se areas may resemble <strong>the</strong> more annual grassland type, Sporobolus (vaginiflorus, neglectus, ozarkanus) - Aristida longispica - Panicum flexile - Panicum capillare Herbaceous Vegetation (CEGL004340) described for Tennessee glades. These shale glades may be so depauperate at this time that <strong>the</strong>y could simply be combined with Quercus stellata - Quercus marilandica / Schizachyrium scoparium - Silphium terebinthinaceum Wooded Herbaceous Vegetation (CEGL005134) (D. Minney pers. comm. 2000). CONSERVATION RANK: G2. The shale glade community has a very restricted distribution and occurrences are small. In Kentucky, where it is most abundant, particularly in <strong>the</strong> Knobs Region, threats are primarily from strip-mining, since oil shales may be strip-mined depending on <strong>the</strong> market. Sites in <strong>Ohio</strong> have been disturbed by agriculture through pasturing (but no clearing), severe erosion, or fire suppression. PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS 310
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PLANT COMMUNITIES OF THE MIDWEST CL
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CONTENTS APPENDIX: PLANT COMMUNITY
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OVERVIEW APPENDIX: PLANT COMMUNITY
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− The USFS (U.S. Forest Service)
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Box 2. State Natural Heritage progr
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INDEX TO ASSOCIATIONS BY ECOLOGICAL
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2.3.4.4. Interior Highlands Shale G
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Acid Peatlands: Northern Shrub/Gram
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Rich Peat Fens: Northeastern Rich F
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Rich Peat Fens: Midwestern Prairie
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Open and Emergent Marshes: Eastern
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Open and Emergent Marshes: Eastern
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