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Ohio subset of Plant Communities of the Midwest ... - NatureServe

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Forests and Woodlands: <strong>Midwest</strong>ern Mesic Hardwood Forests<br />

Fagus grandifolia - Acer saccharum Glaciated <strong>Midwest</strong> Forest<br />

American Beech - Sugar Maple Glaciated <strong>Midwest</strong> Forest<br />

Beech - Maple Glaciated Forest<br />

CEGL005013<br />

DESCRIPTION: This community is characterized by a dense to moderately dense canopy <strong>of</strong> deciduous trees, an<br />

absent to sparse shrub layer, and a moderately to well-developed herbaceous layer. The canopy formed by <strong>the</strong><br />

overstory trees and smaller saplings greatly reduces <strong>the</strong> amount <strong>of</strong> light that reaches lower vegetation strata. This<br />

canopy is composed primarily <strong>of</strong> Acer saccharum and Fagus grandifolia. Liriodendron tulipifera is sometimes<br />

codominant and Carpinus caroliniana, Fraxinus americana, Ostrya virginiana, Quercus rubra, Tilia americana, and<br />

Ulmus americana are <strong>of</strong>ten present in <strong>the</strong> canopy or subcanopy. Diervilla lonicera, Euonymus obovata, Lonicera<br />

canadensis, Sambucus racemosa var. racemosa (= Sambucus pubens), and Viburnum spp. are typical shrubs. The<br />

herbaceous layer is <strong>of</strong>ten diverse, typically including spring ephemerals. Common species include Arisaema<br />

triphyllum (= Arisaema atrorubens), Dryopteris intermedia, Galium aparine, Maian<strong>the</strong>mum canadense (in nor<strong>the</strong>rn<br />

stands), Maian<strong>the</strong>mum racemosum, Osmorhiza claytonii, Podophyllum peltatum, Polygonatum biflorum, Trillium<br />

grandiflorum, and Viola spp.<br />

This community is found on flat to rolling topography (Braun 1950, Pell and Mack 1977). In <strong>the</strong> sou<strong>the</strong>rn parts <strong>of</strong> its<br />

range, it is more likely to be found on steeper slopes than in <strong>the</strong> nor<strong>the</strong>rn portion (Braun 1950). The soils are<br />

predominantly silt loam, loam, or sandy loam and develop over glacial till <strong>of</strong> Wisconsin age (Braun 1950, Dodge and<br />

Harman 1985). This community was found to develop on sites with till 0.4-4.0 m thick in sou<strong>the</strong>rn Michigan (Dodge<br />

and Harman 1985).<br />

COMMENTS: 1, MCS. Wisconsin glaciation history is used to define <strong>the</strong> sou<strong>the</strong>rn limits <strong>of</strong> this type, and fur<strong>the</strong>r work is<br />

needed to determine how floristically distinct <strong>the</strong>se glaciated stands are from sou<strong>the</strong>rn unglaciated stands, which are<br />

placed in <strong>the</strong> Fagus grandifolia - Acer saccharum - Liriodendron tulipifera Unglaciated Forest (CEGL002411).<br />

Liriodendron tulipifera and o<strong>the</strong>r sou<strong>the</strong>rn hardwoods, such as Liquidambar styraciflua, are much less constant in this<br />

type compared to <strong>the</strong> unglaciated stands, as is, perhaps, Asimina triloba. Northward, Betula alleghaniensis and<br />

Betula papyrifera are very infrequent compared to Acer saccharum - Fagus grandifolia - Betula spp. / Maian<strong>the</strong>mum<br />

canadense Forest (CEGL005004). In New York and Pennsylvania this type may cover both <strong>the</strong> glaciated lake till plain<br />

and perhaps some parts <strong>of</strong> <strong>the</strong> glaciated Allegheny Plateau, consistent with its distribution in <strong>Ohio</strong>. Arguing for this is<br />

<strong>the</strong> low dominance or absence <strong>of</strong> Liriodendron tulipifera, low dominance <strong>of</strong> Betula spp., and <strong>the</strong> high dominance <strong>of</strong><br />

Tilia americana. On <strong>the</strong> o<strong>the</strong>r hand, <strong>the</strong> presence <strong>of</strong> Betula lenta, and perhaps that <strong>of</strong> Tsuga canadensis, in that<br />

region are somewhat unusual for this type, indicating <strong>the</strong> more Allegheny influence. And <strong>the</strong> inclusion <strong>of</strong> both<br />

glaciated till plain and plateau habitats in <strong>the</strong> type would broaden <strong>the</strong> environmental parameters. This type may be in<br />

east-central Illinois (Vermillion County, near Danville), but is largely extirpated (B. McClain pers. comm. 1999).<br />

Stands still occur across <strong>the</strong> stateline in Indiana.<br />

CONSERVATION RANK: G3G4. One hundred and fifty-five occurrences have been documented: 3 in Illinois (where <strong>the</strong><br />

community is ranked S4), 86 in Indiana (S3), 36 in Michigan (S3), and 30 in <strong>Ohio</strong> (S3). Although no o<strong>the</strong>r<br />

occurrences have been documented, <strong>the</strong> community is also reported in Ontario (S), and it may occur in New York<br />

(SP). Sizes <strong>of</strong> 129 occurrences total 8895 acres. This type was once a widespread matrix forest that now occupies<br />

small remnants, <strong>of</strong>ten less than 500 acres in size.<br />

DISTRIBUTION: This upland forest community is found in <strong>the</strong> sou<strong>the</strong>rn Great Lakes area <strong>of</strong> <strong>the</strong> United States and<br />

Canada, ranging from Illinois, Indiana, and Michigan, east to Ontario and western New York.<br />

USFS ECOREGIONS: 212Fa:PP, 212Fb:PPP, 221Fa:CCC, 221Fb:CC, 221Fc:CCC, 222De:CCC, 222Ga:CCC, 222Gc:CCC,<br />

222Gd:CCC, 222Ha:CCC, 222Hb:CCC, 222Hc:CCC, 222Hd:CCC, 222Hf:CCC, 222Ia:CC, 222Ib:CC, 222Ic:CC, 222Id:CCP,<br />

222If:CCC, 222Ja:CCC, 222Jb:CCC, 222Jc:CCC, 222Jd:CCC, 222Jg:CCC, 222Jh:CCC, 222Ji:CCC, 222Jj:CCC, 222Kj:CCC,<br />

251De:CCC<br />

CONSERVATION REGIONS: 36:C, 44:C, 45:C, 48:C, 49:C<br />

STATES: IL IN MI NY OH PROVINCES: ON<br />

MIDWEST HERITAGE SYNONYMY: IL mesic upland forest (S) +<br />

IN mesic upland forest +<br />

MI mesic sou<strong>the</strong>rn forest - beech-sugar maple-basswood =<br />

OH beech-sugar maple forest +<br />

OTHER SYNONYMY: Beech - Maple Forest Region (Braun 1950) I, Warren's Woods (Cain 1935) =, Fagus grandifolia -<br />

Acer saccharum - Podophyllum peltatum association (Pell and Mack 1977) =<br />

USNVC HIERARCHY: FAGUS GRANDIFOLIA - ACER SACCHARUM - (LIRIODENDRON TULIPIFERA) FOREST ALLIANCE (I.B.2.N.a)<br />

PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS<br />

421

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