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Ohio subset of Plant Communities of the Midwest ... - NatureServe

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Wooded Swamps and Floodplains: Sou<strong>the</strong>astern Coastal Plain Riverfront and Levee<br />

Bottomland Forests<br />

Betula nigra - Platanus occidentalis Forest<br />

River Birch - Sycamore Forest<br />

River Birch - Sycamore Forest<br />

CEGL002086<br />

DESCRIPTION: Betula nigra and Platanus occidentalis are <strong>the</strong> typical dominants in this type. They are fast-growing<br />

(especially young trees), relatively short-lived, and do not tolerate excessive shade at any stage <strong>of</strong> growth. Betula<br />

nigra prefers acid soils and is <strong>of</strong>ten <strong>the</strong> dominant tree found along streams affected by acid mine drainage. A wide<br />

range <strong>of</strong> canopy species are present because, along with common bottomland species (e.g., Ulmus americana, Acer<br />

rubrum, Acer negundo, Quercus spp., and Celtis laevigata), mesophytic species from nearby uplands and terraces<br />

may be found, including Juglans nigra, Prunus serotina, and Fraxinus americana. A variety <strong>of</strong> swamp species may be<br />

found on more poorly drained sites. Stands are subjected to frequent, fast, short-duration flooding, which contributes<br />

to a thin understory. Commonly encountered herbaceous species include Saururus cernuus, Arisaema dracontium,<br />

Impatiens capensis, Symphyotrichum ontarionis (= Aster ontarionis), and Pilea pumila. This forest harbors a number<br />

<strong>of</strong> ubiquitous species and tends to extend into and mix with adjacent communities, resulting in numerous transitional<br />

variants. Occurrences may have a ra<strong>the</strong>r high percentage <strong>of</strong> standing dead trees. There may be remnant snags from<br />

previous earlier successional communities <strong>of</strong> cottonwood - black willow forests (TNC 1995a).<br />

Betula nigra requires soils near field capacity throughout <strong>the</strong> year but is relatively intolerant <strong>of</strong> flooding. Platanus<br />

occidentalis is also intolerant <strong>of</strong> flooding during <strong>the</strong> growing season and will die if <strong>the</strong> entire tree is inundated for more<br />

than two weeks. The absence <strong>of</strong> this community in <strong>the</strong> lower Mississippi Alluvial Plain is attributed to this intolerance,<br />

as is its most common location on levees <strong>of</strong> smaller rivers. The community usually is found on <strong>the</strong> natural levee <strong>of</strong> <strong>the</strong><br />

watercourse and, <strong>the</strong>refore, is slightly elevated from <strong>the</strong> flats behind <strong>the</strong> levee. The community is more common<br />

along small streams and blackwater streams than along alluvial floodplains, largely because <strong>of</strong> <strong>the</strong> higher sustained<br />

flow rates <strong>of</strong> <strong>the</strong>se larger rivers. It is also found along flowages <strong>of</strong> larger rivers.<br />

This community experiences seasonal inundation or groundwater saturation for approximately 12.5-25% <strong>of</strong> <strong>the</strong> year,<br />

usually during spring and early summer with a frequency <strong>of</strong> 51-100% (<strong>the</strong> Zone IV community <strong>of</strong> Clark and Benforado<br />

1981). Large stream sections with longer flooding durations are more strongly dominated by Betula nigra and Acer<br />

saccharinum alone. The smaller streams on narrow floodplains <strong>of</strong> younger valleys have greater canopy mixtures.<br />

COMMENTS: 2, MCS. As currently described, this association and Betula nigra - Platanus occidentalis / Alnus<br />

serrulata / Boehmeria cylindrica Forest (CEGL007312) appear to be largely overlapping. These two defined<br />

associations need to be reconciled. In Kentucky, this association (CEGL002086) would be found on <strong>the</strong> nor<strong>the</strong>rn part<br />

<strong>of</strong> <strong>the</strong> Daniel Boone National Forest, in contrast to CEGL007312, which is more sou<strong>the</strong>rn. O<strong>the</strong>r bottomland<br />

hardwood communities contain river birch in <strong>the</strong> canopy but at lower densities. It may be helpful to require that Betula<br />

nigra contribute at least 50% (80%) <strong>of</strong> <strong>the</strong> tree density to be placed in this community. The community described<br />

herein is naturally occurring on minimally to undisturbed sites.<br />

CONSERVATION RANK: G5. The community is widespread and common.<br />

DISTRIBUTION: This river birch - sycamore forest community is found throughout <strong>the</strong> eastern United States, ranging<br />

from south-central <strong>Ohio</strong> west to Iowa, south to Arkansas and Mississippi, and east to West Virginia.<br />

USFS ECOREGIONS: 221Ea:CCC, 221Ec:CCC, 221Ed:CCP, 221Ef:CCC, 221Eg:CCC, 221Fa:CCC, 221Fc:CCC, 221Ha:CCC,<br />

221Hb:CCC, 221He:CCC, 222Ad:CCP, 222Ae:CCP, 222Af:CCC, 222Ca:C, 222Ch:C, 222En:CCC, 222Eo:CCC,<br />

222Ga:CCC, 222Ha:CCC, 222If:CCC, 231:P, 232:P, 234:P, 251:, M221Cd:CCC, M222:P, M231:P<br />

CONSERVATION REGIONS: 36:P, 37:P, 39:C, 42:C, 44:C, 45:C, 48:C, 49:C, 50:C<br />

STATES: AR IA IL IN KY MO MS OH OK TN WV PROVINCES: ON<br />

MIDWEST HERITAGE SYNONYMY: IL wet floodplain forest (S) +<br />

IN wet floodplain forest +<br />

MO wet bottomland forest +<br />

OH river birch-maple floodplain forest =<br />

OTHER SYNONYMY: River Birch - Sycamore (61) (Eyre 1980) =, IIA7b. River Birch - Sycamore Riverfront Forest<br />

(Allard 1990) =, Betula nigra - Acer saccharinum / Pilea community (Voigt and Mohlenbrock 1964) =, Acer negundo -<br />

Platanus / Rhus [Toxicodendron] radicans community (Voigt and Mohlenbrock 1964) . Perhaps this type should go<br />

with CEGL007334, a Platanus type, which has not yet been described for Illinois., River Birch Floodplain Forests<br />

(Anderson 1996) =, R13cII4a. Betula nigra - Platanus occidentalis (Foti 1994a)<br />

USNVC HIERARCHY: BETULA NIGRA - (PLATANUS OCCIDENTALIS) TEMPORARILY FLOODED FOREST ALLIANCE (I.B.2.N.d)<br />

PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS<br />

217

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