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Ohio subset of Plant Communities of the Midwest ... - NatureServe

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Wooded Swamps and Floodplains: <strong>Midwest</strong>ern Wet Flatwoods<br />

Fagus grandifolia - Quercus alba - (Quercus michauxii) - Acer rubrum Flatwoods Forest<br />

American Beech - White Oak - (Swamp Chestnut Oak) - Red Maple Flatwoods Forest<br />

Beech - Oak - Red Maple Flatwoods<br />

CEGL005015<br />

DESCRIPTION: This community has a moderately dense to dense canopy <strong>of</strong> deciduous trees. Fagus grandifolia is<br />

consistently common. O<strong>the</strong>r common trees include Acer rubrum, Liriodendron tulipifera, Nyssa sylvatica, Quercus<br />

alba, Quercus bicolor, Quercus palustris, and Quercus michauxii. Fraxinus americana, Liquidambar styraciflua, and<br />

Quercus shumardii may be present. Asimina triloba, Carpinus caroliniana, Lindera benzoin, Toxicodendron radicans,<br />

and Sambucus canadensis are common shrubs. Vitis labrusca is a common, and perhaps characteristic, vine.<br />

Abundant understory species include <strong>the</strong> ferns Botrychium oneidense and Onoclea sensibilis and <strong>the</strong> herbs<br />

Chimaphila maculata, Cinna arundinacea, Impatiens capensis (= Impatiens biflora), Panax trifolius, Podophyllum<br />

peltatum, and Rubus hispidus.<br />

This community is usually found on loess-covered glacial drift <strong>of</strong> Illinoisan age, with level topography and poorly<br />

drained silt loam soils (Braun 1936). The substrate can form a mosaic <strong>of</strong> slightly higher, wet-mesic raised areas, and<br />

wet depressions. An impermeable layer beneath <strong>the</strong> topsoil slows drainage <strong>of</strong>ten resulting in wet soil conditions<br />

(Chapman 1942). In droughty periods, <strong>the</strong> upper soils dry out, and <strong>the</strong> impermeable subsoil limits <strong>the</strong> availability <strong>of</strong><br />

deeper water sources.<br />

COMMENTS: 2, MCS. This type is based largely on <strong>the</strong> Indiana state type - Bluegrass Till Plain Flatwoods (Homoya et<br />

al. 1988) and <strong>Ohio</strong>'s Wet Beech Forests (Anderson 1996). The type consists <strong>of</strong> a mosaic <strong>of</strong> raised, wet-mesic areas<br />

and depressions that are wet. The vegetation varies accordingly. Floristic distinctions between <strong>the</strong> bluegrass region<br />

and <strong>the</strong> central tillplain need to be worked out, but <strong>the</strong> bluegrass occurrences do contain Quercus michauxii and<br />

Liquidambar styraciflua. Anderson (1996) cites Braun (1936) as noting that Carpinus caroliniana, Photinia floribunda<br />

(= Aronia prunifolia), Spiraea tomentosa, Ilex verticillata, Vitis labrusca, and Viburnum dentatum var. deamii are<br />

characteristic <strong>of</strong> this type. To help define <strong>the</strong> type, stands may require at least 25% basal area or cover <strong>of</strong> beech<br />

(Anderson 1996). In Indiana, this type may also occur in <strong>the</strong> southwestern part <strong>of</strong> <strong>the</strong> state (Spencer County).<br />

CONSERVATION RANK: G3. There are probably fewer than 100 occurrences rangewide. Seven have been<br />

documented in <strong>Ohio</strong>, where <strong>the</strong> community is ranked S3, and 11 in Indiana (S2). The community occurs in 6<br />

ecoregional subsections. There are probably fewer than 10,000 acres rangewide. Sizes <strong>of</strong> 11 occurrences total 1480<br />

acres. The community occurs on level, poorly drained silt loams (usually loess-covered glacial drift) with an<br />

impenetrable layer beneath <strong>the</strong> topsoil that impedes drainage and accessibility to deeper water.<br />

DISTRIBUTION: This lowland forest community is found in <strong>the</strong> United States in sou<strong>the</strong>astern Indiana and<br />

southwestern <strong>Ohio</strong>.<br />

USFS ECOREGIONS: 222Ek:CCC, 222Fc:CC, 222Fd:CCC, 222Fe:CCC, 222Ff:CCC, 222Ha:CCC, 222Hc:CCC<br />

CONSERVATION REGIONS: 44:C, 45:C, 48:C, 49:C<br />

STATES: IN OH PROVINCES:<br />

MIDWEST HERITAGE SYNONYMY: IN bluegrass till plain flatwoods =<br />

OH beech-oak-red maple forest =<br />

OTHER SYNONYMY: Beech - White Oak - Sweet Gum Community (Braun 1936) F, Beech - White Oak - Red Maple<br />

Community (Braun 1936) F, Beech - White Oak - Red Maple - Sweet Gum Community (Braun 1936) F, Beech -<br />

White Oak Associes I, Wet Beech Forests (Anderson 1996) B. Anderson describes differences between <strong>the</strong> stands<br />

in southwest <strong>Ohio</strong> from those in <strong>the</strong> west/northwest part <strong>of</strong> <strong>the</strong> state. Parts <strong>of</strong> Anderson's mixed swamp type may also<br />

fall into this type, depending on <strong>the</strong> percentage <strong>of</strong> beech used to define his wet beech forest type.<br />

USNVC HIERARCHY: FAGUS GRANDIFOLIA - QUERCUS SPP. - ACER SPP. FOREST ALLIANCE (I.B.2.N.a)<br />

PLANT COMMUNITIES OF THE MIDWEST – 2001. APPENDIX: ASSOCIATION DESCRIPTIONS<br />

188

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