THE UNIVERSITY OF CALGARY Eric Snively A ... - Ohio University

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Figure 3.4. Freedom of intermetatarsal movement, as revealed by CT scans of a right metatarsus of the tyrannosaurid Gorgosaurus libratus (94.1 2.602). Arrows show a sliding motion evident between MT II and MT III, and a slight rotational motion between MT IV and MT III.
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THE UNIVERSITY OF CALGARY Functiona
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Many figures in this thesis use col
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ACKNOWLEûGEM ENTS Many friends and
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Approval page Acknowledgements Tabi
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CONCLUDING COMMENTS Figures for Cha
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CHAPTER 5: Mechanical and phylogene
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LIST OF FIGURES Figure 1 .l. Phylog
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Figure 3.6. Osteological correlates
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AMNH GI HMN IVPP MOR NMC OMNH PIN P
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and constructional morphology revea
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1985, Bryant and Seymour 1990). If
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surfaces of a Tyrannosaurus rex spe
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3. Troodontidae (Figure 1.2): Trwdo
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Hypothesized functions of the tyran
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1. Atomization (Chapters 2 and 3).
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Figure 1.1. P hylogenetic diagram o
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Figure 1.2. Phylogenetic diagram of
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Figure 1 .S. Skeletal restorations
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did not quantify the degree of prox
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qualitative debate over arctometata
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This hypothesis focuses on tyrannos
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Table 2.1. Theropod and Plafeosauru
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specimens in the figures reflects a
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were not attempted. While this limi
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Chapter 3). 1 now address variation
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- Ç6'P 1 68'€tr 1 L'OC 99'66 L'9
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Table 2.3: Log-transformed values f
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experiences a sharp laterally indin
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ecause it can be cursorily dassifie
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arctometatarsalian specimens, the l
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) Shaf€ and region of proximal ar
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a) Ginglymus. Figure 2.14 shows the
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) Shaft and region of proximal arti
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measurements of width pV2S%TU-DE an
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Table 2.4b Variancelmeasurement eac
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the largest sum of linear measureme
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Table 2.5: Specimen statistics Herr
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and Sinraptor dongi specimens. With
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In addition, a hook like proximal a
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oth physical narrowing and elongati
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1s the MT 111 shape segregation fun
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tyrannosaurids, ornithomimid, and t
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proximal constriction, white EImisa
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Figure 2.2. Tyrannosaurid, trwdonti
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Figure 2.4. Dromaeosaurid and carno
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- Anterior (flexor) surface medial
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Figure 2.7. Left MT III of Tyrannos
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Figure 2.8. Left MT III of Gorgosau
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Figure 2.9. Left MT III of an omith
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Figure 2.10. Left metatarsus of Tmd
Page 113 and 114: Figure 2.1 1. Left MT III of Elmisa
Page 115 and 116: Figure 2.12. Right MT III of Deinon
Page 117 and 118: Figure 2.13. Left MT III of Allosau
Page 119 and 120: Figure 2.14. Left metatarsus of Sin
Page 121 and 122: Figure 2.15. Plots of MT Ill specir
Page 123: Figure 2.16. Plot of third metatars
Page 126 and 127: tv . ... . Ri. . . .
Page 129 and 130: Figure 2.19. Plot of third metatars
Page 131 and 132: CHAPTER 3: Tensile keystone model o
Page 133 and 134: dynamically transmitted Iommotor fo
Page 135 and 136: Table 3.1 . Metatarsi examined in t
Page 137 and 138: interrnetatarsal dynamics in these
Page 139 and 140: compelled analysis of movement evid
Page 141 and 142: Once prepared, TMP 94.1 2.602 and i
Page 143 and 144: Daspletosaurus tomsus (MOR 590), an
Page 145 and 146: 2. Freedom of movement inferred fro
Page 147 and 148: Table 3.2. Surface areas of intemet
Page 149 and 150: phylogenetic bracketing Wtmer 1995)
Page 151 and 152: 4) Forces from anterior displacerne
Page 153 and 154: e expected for tyrannosaurid interm
Page 155 and 156: The preceding discussion derives fr
Page 157 and 158: weapons sparring), and the arctomet
Page 159: Figure 3.1. Schematic representatio
Page 163: Figure 3.3. CT reconstruction of ri
Page 169: Figure 3.6. Osteological correlates
Page 173: Figure 3.8. Osteological correlates
Page 177: Figure 3.10. Step sequence of Gorgo
Page 185: Figure 3.12. CT reconstructions of
Page 189: Figure 3.14. Torsional loading tran
Page 192 and 193: Lu- dorsal 81igarnents
Page 195 and 196: CHAPTER 4: Finite element model of
Page 197 and 198: forces are applied to nodes on the
Page 199 and 200: Material and Ioading regimes of fhe
Page 201 and 202: Two analyses were fun in order to t
Page 203 and 204: Dunng Iinear progression when an an
Page 205 and 206: This quantity approximates the vert
Page 207 and 208: to its right foot to prepare for a
Page 209 and 210: elements in the finite element rnes
Page 211 and 212: Table 4.1. Material properties of b
Page 213 and 214: Photoshop 3 for Macintosh, to ensur
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this program. NUAGES typically foun
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1) Informative strain results versu
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of strain energy evident in Figure
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Specifically, osteological correlat
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custiioned the rnetatarsals against
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Figure 4.2. Initial loads and bound
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Figure 4.4. Simplified representati
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Figure 4.6. Strain distribution in
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The analyses of the Gorgosaurus Iib
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individual organs with ideally effi
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These considerations of ligament mo
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The precise scaling of metatarsal l
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tenable from the available evidence
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equivalent horsepower, 15 economy c
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Other aptations refiect multiple fu
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This and other systernatically info
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(Russell 1975, McGregor 2000). Meta
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was the default developmental patte
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Arnong dromaeosaurids and birds, th
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hypothesis. Fossil evidence of basa
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suggestive of grasping ability. Neo
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available evidence without propriet
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Figure 5.1. Phylogeny of the Therop
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LITERTURE ClTED Alexander R.M. 1977
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Carter D.R., Fyhrie, D.P., and Whal
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Galton P.M. 1982. Elaphrosaurus, an
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Hutchinson J.R., Stidham T.A., Sniv
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Novas F.E. 1993. New information on
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Stein B.R. and Casinos 1997. What i
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APPENDIX Principal Components Analy
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highest degree, and so on. Specimen
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THE UNIVERSITY OF CALGARY Eric Snively A ... - Ohio University

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