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Examples where Art. 40.2 does not apply:<br />

Suter (1909) placed Columbella Lamarck,<br />

1 799, and Pyrene Röding, 1 798, in the same<br />

family. He did not treat them as synonyms<br />

but, because Pyrene was the senior name,<br />

he used the new name Pyrenidae instead of<br />

CoLUMBELLiDAE Swainson, 1840. Pyrenidae is<br />

not a replacement name in the sense of Art.<br />

40.2, and it does not take the precedence of<br />

COLUMBELLIDAE.<br />

Dall (1866) established Pompholiginae based<br />

on Pompholyx Lea, 1856. However, the type<br />

genus is a junior homonym of Pompholyx<br />

Gosse, 1851 [Rotifera]. Lindholm (1927b)<br />

replaced Pompholyx and Pompholiginae with<br />

the names Pompholycodea and Pompholycodeinae<br />

respectively. The replacement was<br />

not a consequence of synonymy of the type<br />

genus and Art. 40.2 does not apply.<br />

Examples where Art. 40.2 applies:<br />

Suter (1913) placed Dolium Lamarck, 1 801<br />

in synonymy of Tonna Brünnich, 1772, and<br />

replaced Doliidae Latreille, 1825, with the<br />

new name Tonnidae. Tonnidae is in prevail-<br />

ing usage and is to be maintained, with the<br />

precedence of Doliidae. It should be cited<br />

Tonnidae Suter, 1913(1825).<br />

Beyond such cases that fit literally to the<br />

wording of the Code, there is a broader ar-<br />

NOMENCLÁTOR OF GASTROPOD FAMILIES 11<br />

ray of cases in which the author establish-<br />

ing the younger family-group name did not<br />

explicitly state that he did so "because of the<br />

synonymy of the type genus".<br />

For instance, when he established the name<br />

DisciNAE, Thiele (1931 [in 1929-1935]) did<br />

not state that he was replacing Patulinae<br />

Tryon, 1866, because of the synonymy of<br />

Patula Held, 1 837, nor did he even mention<br />

the name Patulinae, but he cited Patula as a<br />

synonym oWiscus Fitzinger, 1833. We have<br />

treated this as a situation covered by Art.<br />

40.2. DisciDAE is in prevailing usage and is<br />

to be maintained, with the precedence of<br />

Patulinae. It should be cited Discidae Thiele,<br />

1931 (1866).<br />

Departing still a little further from the letter<br />

of Art. 40.2, there are cases in which the<br />

author establishing the younger family-group<br />

name not only did not explicitly stated that<br />

he was doing so "because of the synonymy<br />

of the type genus" but not even mentioned<br />

the synonymy of the genera involved.<br />

For instance, when he established<br />

Melampidae, Stimpson (1851) did not state<br />

he was replacing Conovulidae W. Clark,<br />

1850, because of the synonymy of<br />

Conovulus Bowdich, 1822, nor did he mentioned<br />

the names Conovulidae or Conovulus.<br />

However, Melampus Montfort, 1810, and<br />

Conovulus are (objective) synonyms, and<br />

Melampinae is in prevailing usage. We have<br />

also treated this as a situation covered by<br />

Art. 40.2, and we have maintained<br />

Melampinae Stimpson, 1851 (1850), as the<br />

valid name.<br />

Names that are invalid under Art. 39, or because<br />

they have been placed on the Official<br />

Index, are permanently invalid, and<br />

cannot be used as valid in any classifica-<br />

tion. Taxonomical synonyms are also invalid,<br />

but only within the frame of a classification,<br />

and these may be resurrected by another<br />

author who has a different opinion about<br />

classification.<br />

Example:<br />

Our classification recognizes a family<br />

Phenacolepadidae with three synonyms, two<br />

of which are invalid under Art. 39.<br />

Family Phenacolepadidae Pilsbry, 1895<br />

[= Scutellidae Angas, 1871 (inv.);<br />

= Scutellinidae Dall, 1889 (inv.);<br />

= Shinkailepadidae Okutani, Saito &<br />

Hashimoto, 1989]<br />

A hypothetical author considering that the<br />

family necessitates more ranks between<br />

family and genus could come with another<br />

classification, e.g.:<br />

Family Phenacolepadidae Pilsbry, 1895<br />

SF Phenacolepadinae Pilsbry, 1895<br />

[= Scutellidae Angas, 1871 (inv.);<br />

= Scutellinidae Dall, 1889 (inv.)]<br />

SF Shinkailepadinae Okutani, Saito &<br />

Hashimoto, 1989<br />

Principle of Coordination<br />

Article 36 states that: "A name established<br />

for a taxon at any rank in the family group is<br />

deemed to be simultaneously established with<br />

the same author and date for taxa based upon<br />

the same name-bearing type (type genus) at<br />

other ranks in the family group, with appropri-<br />

ate mandatory change of suffix".<br />

Example:<br />

Ellis (1926) established the name Milacidae<br />

at family rank. He is deemed to have established<br />

that name at any other rank in the fam-<br />

ily group. The author and date of Milacinae<br />

is Ellis, 1926, despite that it was declared a<br />

new subfamily by Germain (1931).

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