A monograph of the British fossil corals - kreidefossilien.de
A monograph of the British fossil corals - kreidefossilien.de
A monograph of the British fossil corals - kreidefossilien.de
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iv<br />
BRITISH FOSSIL CORALS.<br />
In cases <strong>of</strong> this kind <strong>the</strong> distinction between <strong>the</strong> Polypi and <strong>the</strong> Bryozoa is always<br />
ren<strong>de</strong>red easy by <strong>the</strong> most superficial examination <strong>of</strong> <strong>the</strong> s<strong>of</strong>t parts <strong>of</strong> <strong>the</strong> animal ; but it is<br />
sometimes a matter <strong>of</strong> great difficulty for <strong>the</strong> palaeontologist, who is<br />
necessarily <strong>de</strong>prived <strong>of</strong><br />
all such resources, and can only be gui<strong>de</strong>d by <strong>the</strong> peculiarities observable in <strong>the</strong> ossified<br />
tissues.<br />
In general, <strong>the</strong> distinction between Corals and Spongidae is also very easy, for <strong>the</strong><br />
lamellar structure, so prevalent among <strong>the</strong> former, is never met with in <strong>the</strong> latter ; but in<br />
some Polypidoms (certain Milleporidse for example), <strong>the</strong> vertical plates disappear, and <strong>the</strong><br />
mural tissue becomes extremely porous, irregulai', and abundant, so as to resemble much<br />
<strong>the</strong> reticulated mass formed by <strong>the</strong> stony skeleton <strong>of</strong> some Spongidae, where <strong>the</strong> oscula<br />
and aquiferous canals are on <strong>the</strong> contrary more regular than usual. In cases <strong>of</strong> this kind<br />
it may be necessary to seek for distinctive characters in <strong>the</strong> internal structure <strong>of</strong> <strong>the</strong><br />
Zoophyte ; and, in<strong>de</strong>pen<strong>de</strong>ntly <strong>of</strong> <strong>the</strong> benefit to be obtained by <strong>the</strong> microscopical investigation<br />
<strong>of</strong> <strong>the</strong> tissue itself, it will sometimes be found useful to examine <strong>the</strong> form <strong>of</strong> <strong>the</strong><br />
tubular cavities which perva<strong>de</strong> <strong>the</strong> mass, and correspond ei<strong>the</strong>r to <strong>the</strong> visceral chambers <strong>of</strong><br />
<strong>the</strong> Polypi, or to <strong>the</strong> great aquiferous ducts <strong>of</strong> <strong>the</strong> Spongida; ; for in <strong>the</strong> first instance<br />
<strong>the</strong>y are always simple, Avhereas in <strong>the</strong> latter <strong>the</strong>y are more or less ramified.<br />
§IL<br />
The external forms <strong>of</strong> Corals vary consi<strong>de</strong>rably, but are in general more <strong>de</strong>pen<strong>de</strong>nt on<br />
<strong>the</strong> mo<strong>de</strong> <strong>of</strong> aggregation <strong>of</strong> <strong>the</strong> different individuals produced by a common parent than<br />
on <strong>the</strong> mo<strong>de</strong> <strong>of</strong> organization pecuhar to<br />
<strong>the</strong> animals to which <strong>the</strong>se tegumentary skeletons<br />
belong. Characters <strong>de</strong>rived from <strong>the</strong>se forms can <strong>the</strong>refore be but <strong>of</strong> little avail for <strong>the</strong><br />
natm-al arrangement <strong>of</strong> Polypi ; and <strong>the</strong> classification <strong>of</strong> <strong>the</strong>se Zoophytes, like that <strong>of</strong> <strong>the</strong><br />
higher animals, must be foun<strong>de</strong>d on <strong>the</strong> principal modifications observable in <strong>the</strong>ir structm-e.<br />
It would lead us too far from <strong>the</strong> special object <strong>of</strong> this Monograph, if we were to<br />
enter on <strong>the</strong> investigation <strong>of</strong> <strong>the</strong> anatomical facts which alone can furnish satisfactory<br />
elements for such a classification ; but in or<strong>de</strong>r to facilitate <strong>the</strong> study <strong>of</strong> <strong>the</strong> Corals about<br />
to be <strong>de</strong>scribed, it may be useful for us to revert to a few <strong>of</strong> <strong>the</strong> leading points in <strong>the</strong><br />
structure <strong>of</strong> Polyi^i, and to <strong>de</strong>fine some <strong>of</strong> <strong>the</strong> expressions which we shall <strong>of</strong>ten have to<br />
employ.'<br />
The ScLERENCHYMA, or har<strong>de</strong>ned tissue <strong>of</strong> Polypi, by which Corals are formed, is<br />
always a portion <strong>of</strong> <strong>the</strong> tegumentary system <strong>of</strong> <strong>the</strong>se Zoophytes, but, as we have abeady<br />
stated, it may be produced in two very different ways. In some cases it is <strong>the</strong> result <strong>of</strong><br />
a sort <strong>of</strong> ossification <strong>of</strong> <strong>the</strong> chorion or principal tunic <strong>of</strong> <strong>the</strong> Polypi ; in o<strong>the</strong>rs it grows on<br />
'<br />
For more ample <strong>de</strong>tails on this subject we must refer to our " Memoir on <strong>the</strong> Structure and Development<br />
<strong>of</strong> Corals," published in <strong>the</strong> Annales <strong>de</strong>s Sciences Naturelles, 3"° serie, t. ix.