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13th International Conference on Membrane Computing - MTA Sztaki

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P. Ramón, A. Troina<br />

Fig. 1. Metapopulati<strong>on</strong> dynamics.<br />

describes a selective pressure driving populati<strong>on</strong>s evoluti<strong>on</strong> through the logistic<br />

model [47]:<br />

dN<br />

(1<br />

dt = r · N · − N )<br />

K<br />

where N represents the number of individuals in the populati<strong>on</strong>.<br />

CWC Modelling 3 (Logistic Model) The logistic model with growth rate r<br />

and carrying capacity K, for an envir<strong>on</strong>ment modelled by a compartment with<br />

label l, can be encoded within CWC using two stochastic rewrite rules describing<br />

(i) a reproducti<strong>on</strong> event for a single individual at the given rate and (ii) a<br />

death event modelled by a fight between two individuals at a rate that is inversely<br />

proporti<strong>on</strong>al to the carrying capacity:<br />

r<br />

l : a ↦−→ a a<br />

2·r<br />

K−1<br />

l : a a ↦−→ a<br />

If N is the number of individuals of species a, the number of possible reactants<br />

for the first rule is N and the number of possible reactants for the sec<strong>on</strong>d rule<br />

is, in the exact stochastic model, ( )<br />

N<br />

2 =<br />

N·(N−1)<br />

2<br />

, i.e. the number of distinct<br />

pairs of individuals of species a. Multiplying this values by the respective rates<br />

we get the propensities of the two rules and can compute the value of N when<br />

the equilibrium is reached (i.e., when the propensities of the two rules are equal):<br />

r · N = 2·r<br />

K−1 · N·(N−1)<br />

2<br />

, that is when N = 0 or N = K.<br />

For a given species, this model allows to describe different growth rates and<br />

carrying capacities in different ecological regi<strong>on</strong>s. Identifying a CWC compartment<br />

type (through its label) with an ecological regi<strong>on</strong>, we can define rules<br />

describing the growth rate and carrying capacity for each regi<strong>on</strong> of interest.<br />

392

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