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13th International Conference on Membrane Computing - MTA Sztaki

13th International Conference on Membrane Computing - MTA Sztaki

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P. Ramón, A. Troina<br />

Fig. 3. Competiti<strong>on</strong> and Mutualism.<br />

(“who eats who”) in an ecological community [30]. At the base of the food<br />

web there are autotroph species 12 , also called basal species. A food chain is a<br />

linear feeding pathway that links m<strong>on</strong>ophagous c<strong>on</strong>sumers (with <strong>on</strong>ly <strong>on</strong>e exiting<br />

trophic link) from a top c<strong>on</strong>sumer, usually a larger predator, to a basal species.<br />

The length of a chain is given by the number of links between the top c<strong>on</strong>sumer<br />

and the base of the web. The influence that the elements of a food web have<br />

<strong>on</strong> each other determine important features of an ecosystem like the presence<br />

of str<strong>on</strong>g interactors (or keyst<strong>on</strong>e species), the total number of species, and the<br />

structure, functi<strong>on</strong>ality and stability of the ecological community.<br />

To model quantitatively a trophic link between species a and b (i.e., a particular<br />

kind of competiti<strong>on</strong>) we might use Lotka-Volterra equati<strong>on</strong>s [48]:<br />

dN b<br />

dt<br />

= N b · (r b − α · N a )<br />

dN a<br />

dt<br />

= N a · (β · N b − d)<br />

where N a and N b are the numbers of predators and preys, respectively, r b is the<br />

rate for prey growth, α is the prey mortality rate for per-capita predati<strong>on</strong>, β<br />

models the efficiency of c<strong>on</strong>versi<strong>on</strong> from prey to predator and d is the mortality<br />

rate for predators.<br />

CWC Modelling 5 (Trophic Links) Within a compartment of type l, given<br />

a predati<strong>on</strong> mortality α and c<strong>on</strong>versi<strong>on</strong> from prey to predator β, we can encode<br />

in CWC a trophic link between individuals of species a (predator) and b (prey)<br />

by the following rules:<br />

l : a b ↦−→ α<br />

a<br />

l : a b ↦−→ β<br />

a a b<br />

12 Self-feeding: able to produce complex organic compounds from simple inorganic<br />

molecules and light (by photosynthesis) or inorganic chemical reacti<strong>on</strong>s (chemosynthesis).<br />

396

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