Dioscorea quaternata, Eupatorium purpureum, Galium latifolium, Galax urceolata, Goodyera pubescens, Houstonia purpurea var.purpurea, Lysimachia quadrifolia, Maianthemum racemosum ssp. racemosum, Medeola virginiana, Polygonatum biflorum,Polystichum acrostichoides, Solidago curtisii (= Solidago caesia var. curtisii), and Uvularia puberula. Common vines are Smilaxrotundifolia, Smilax glauca, and Vitis aestivalis. This forest is distinguished from High Elevation Red Oak forests [see associations inI.B.2.N.a Quercus rubra Montane <strong>Forest</strong> Alliance (A.272)] by lack of species such as Betula alleghaniensis, Ilex montana, Vacciniumsimulatum, and by lacking abundant Hamamelis virginiana, as well as its occurrence at lower elevations. In the Southern Blue RidgeEscarpment region, these montane oak - hickory forests seem to occupy environments intermediate between more protected forestsdominated by Quercus alba and drier, more exposed Quercus prinus forests.Environment: This association includes Quercus rubra forests at intermediate elevations (mostly below 3500 feet, ranging from2000-4000 feet) in the Southern Blue Ridge Escarpment, and may possibly range into adjacent areas of the Central Appalachians andCumberland Plateau. These forests occur on mostly northern to eastern and southeastern, mid to upper, moderately steep slopes ofintermediate exposure over acidic soils.Vegetation: The canopy is dominated by Quercus rubra, often with Acer rubrum and/or Liriodendron tulipifera codominating, andoccasionally with a high component of Quercus alba in the canopy. Other minor canopy species may include Betula lenta, Caryaalba, Carya glabra, Halesia tetraptera, Quercus prinus, and Magnolia fraseri. The subcanopy and sapling strata include the canopyspecies, as well as Halesia tetraptera, Betula lenta, Tsuga canadensis, Cornus florida, Acer pensylvanicum, and Oxydendrumarboreum. The shrub stratum is typically sparse but may have local dominance by Gaylussacia ursina or Rhododendron maximum.Other typical species in the shrub stratum include Castanea dentata, Calycanthus floridus, Pyrularia pubera, Rhododendroncalendulaceum, Vaccinium corymbosum, and Viburnum acerifolium. Herbaceous cover is sparse to moderate but can be species rich.Ferns can be locally dominant, typically Thelypteris noveboracensis and Athyrium filix-femina ssp. asplenioides. Other typical speciesinclude Eurybia divaricata (= Aster divaricatus), Carex spp. (e.g., Carex aestivalis, Carex debilis, Carex digitalis, Carex laxifloravar. laxiflora, Carex pensylvanica), Chimaphila maculata (= var. maculata), Desmodium nudiflorum, Dioscorea quaternata,Eupatorium purpureum, Galium latifolium, Galax urceolata, Goodyera pubescens, Houstonia purpurea var. purpurea, Lysimachiaquadrifolia, Maianthemum racemosum ssp. racemosum, Medeola virginiana, Polygonatum biflorum, Polystichum acrostichoides,Solidago curtisii (= Solidago caesia var. curtisii), and Uvularia puberula. Common vines are Smilax rotundifolia, Smilax glauca, andVitis aestivalis. This forest is distinguished from High Elevation Red Oak forests [see associations in I.B.2.N.a Quercus rubraMontane <strong>Forest</strong> Alliance (A.272)] by lack of species such as Betula alleghaniensis, Ilex montana, Vaccinium simulatum, and bylacking abundant Hamamelis virginiana, as well as its occurrence at lower elevations. In the Southern Blue Ridge Escarpment region,these montane oak - hickory forests seem to occupy environments intermediate between more protected forests dominated by Quercusalba and drier, more exposed Quercus prinus forests.Dynamics: No informationSimilar Associations:• Quercus alba - Quercus (rubra, prinus) / Rhododendron calendulaceum - Kalmia latifolia - (Gaylussacia ursina) <strong>Forest</strong>(CEGL007230) -- contains more than 50% Quercus alba in the canopy.• Quercus rubra / (Vaccinium simulatum, Rhododendron calendulaceum) / (Dennstaedtia punctilobula, Thelypterisnoveboracensis) <strong>Forest</strong> (CEGL007300) -- is a high-elevation forest.Related Concepts:• IA6h. Montane Oak - Hickory <strong>Forest</strong> (Allard 1990) B• Montane Red Oak-Hickory <strong>Forest</strong> (Schafale 1998b) ?• Oak - Chestnut - Hickory <strong>Forest</strong> (Ambrose 1990a) BClassification Comments: This association was originally defined from the Chattooga Basin Project (S. Simon pers. comm.) andlater refined with information from the Great Smoky Mountains. Global name and concept may need revision as more informationbecomes available. This association may be a subset of the more broadly defined Quercus alba - Quercus (rubra, prinus) /Rhododendron calendulaceum - Kalmia latifolia - (Gaylussacia ursina) <strong>Forest</strong> (CEGL007230) but is distinguished by the dominanceof Quercus rubra, generally protected topographic setting, and may represent areas formerly dominated by Quercus rubra andCastanea dentata. This type replaced Castanea dentata in Virginia (G. Fleming pers. comm.).CONSERVATION RANKING & RARE SPECIESGRank: G4? (2000-1-3): This community is uncommon but secure within its range. It is often overlooked in surveys or notrecognized as distinct, thus it is much more common than the number of documented occurrences suggests. Resolution of taxonomicissues that distinguish this community from similar associations may lead to a range extension.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This association is found in the Southern Blue Ridge Escarpment and may possibly range into adjacent areas of the CentralAppalachians and Cumberland Plateau.Subnations: GA, NC, SC, TN, VA?TNC Ecoregions: 50:?, 51:C, 59:?USFS Ecoregions: M221Dc:CCP, M221Dd:CCCFederal Lands: NPS (Carl Sandburg Home, Great Smoky Mountains); USFS (Chattahoochee, Nantahala, <strong>Sumter</strong>)Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 200478Copyright © 2004 <strong>NatureServe</strong>
ELEMENT SOURCESReferences: Allard 1990, Ambrose 1990a, Nelson 1986, Peet et al. unpubl. data 2002, Schafale 1998b, Schafale and Weakley 1990,Simon pers. comm., Southeastern Ecology Working Group n.d.ROCK CHESTNUT OAK - (NORTHERN RED OAK) - HICKORY SPECIES / SOURWOOD - FLOWERING DOGWOODFORESTELEMENT IDENTIFIERSNVC association: Quercus prinus - (Quercus rubra) - Carya spp. / Oxydendrum arboreum - Cornus florida <strong>Forest</strong>Database Code: CEGL007267Formation: Lowland or submontane cold-deciduous forest (I.B.2.N.a)Alliance: Quercus prinus - Quercus rubra <strong>Forest</strong> Alliance (A.250)ELEMENT CONCEPTSummary: This community is known from low to intermediate elevations of the Southern Blue Ridge escarpment and Piedmonttransition areas. It occurs on relatively exposed landforms below 3000 feet elevation (1200-2900 feet), on moderately steep to steep,convex, middle to upper slopes and ridges, with mostly northern to southwestern aspects. Canopies are dominated by Quercus prinus,with Acer rubrum often codominating. Other species that can have significant canopy coverage include Carya glabra, Liriodendrontulipifera, and Quercus rubra. The subcanopy is commonly dominated by Cornus florida. Additional canopy and subcanopy speciescan include Quercus velutina, Carya alba, Halesia tetraptera var. monticola, Nyssa sylvatica, Robinia pseudoacacia, Magnoliafraseri, and Oxydendrum arboreum. The shrub stratum is sparse with no clear dominant. Some typical shrub species includeGaylussacia ursina, Hydrangea arborescens, Hydrangea radiata, Kalmia latifolia, Magnolia fraseri, Sassafras albidum, and,Vaccinium pallidum. Common vines are Smilax rotundifolia, Smilax glauca, Vitis aestivalis, Vitis rotundifolia, and Vitis vulpina. Herbcover is sparse, but diversity and species composition vary among occurrences. Some of the more typical species include Eurybiadivaricata (= Aster divaricatus), Chimaphila maculata, Desmodium nudiflorum, Dichanthelium spp. (e.g., Dichanthelium boscii,Dichanthelium commutatum, Dichanthelium dichotomum), Dioscorea quaternata, Galium latifolium, Houstonia purpurea,Lysimachia quadrifolia, Maianthemum racemosum ssp. racemosum, Polystichum acrostichoides, Prenanthes spp., Thalictrumthalictroides, Thelypteris noveboracensis, Uvularia perfoliata, Uvularia puberula, Uvularia sessilifolia, and Viola spp. (e.g., Violablanda, Viola hastata, Viola X palmata, Viola tripartita). Some occurrences may have areas of exposed rock.Environment: This community is known from low to intermediate elevations of the Southern Blue Ridge escarpment and Piedmonttransition areas. It occurs on relatively exposed landforms below 3000 feet elevation (1200-2900 feet), on moderately steep to steep,convex, middle to upper slopes and ridges, with mostly northern to southwestern aspects.Vegetation: The canopies of stands of this type are dominated by Quercus prinus, with Acer rubrum often codominating. Otherspecies that can have significant canopy coverage include Carya glabra, Liriodendron tulipifera, and Quercus rubra. The subcanopyis commonly dominated by Cornus florida. Additional canopy and subcanopy species can include Quercus velutina, Carya alba,Halesia tetraptera var. monticola, Nyssa sylvatica, Robinia pseudoacacia, Magnolia fraseri, and Oxydendrum arboreum. The shrubstratum is sparse with no clear dominant. Some typical shrub species include Gaylussacia ursina, Hydrangea arborescens,Hydrangea radiata, Kalmia latifolia, Magnolia fraseri, Sassafras albidum, and, Vaccinium pallidum. Common vines are Smilaxrotundifolia, Smilax glauca, Vitis aestivalis, Vitis rotundifolia, and Vitis vulpina. Herb cover is sparse, but diversity and speciescomposition vary among occurrences. Some of the more typical species include Eurybia divaricata (= Aster divaricatus), Chimaphilamaculata, Desmodium nudiflorum, Dichanthelium spp. (e.g., Dichanthelium boscii, Dichanthelium commutatum, Dichantheliumdichotomum), Dioscorea quaternata, Galium latifolium, Houstonia purpurea, Lysimachia quadrifolia, Maianthemum racemosum ssp.racemosum, Polystichum acrostichoides, Prenanthes spp., Thalictrum thalictroides, Thelypteris noveboracensis, Uvularia perfoliata,Uvularia puberula, Uvularia sessilifolia, and Viola spp. (e.g., Viola blanda, Viola hastata, Viola X palmata, Viola tripartita).Dynamics: No informationSimilar Associations:• Quercus prinus - Quercus rubra / Hamamelis virginiana <strong>Forest</strong> (CEGL006057)Related Concepts:• Chestnut Oak <strong>Forest</strong> (Herb Subtype) (Schafale 1998b) ?• IA6h. Montane Oak - Hickory <strong>Forest</strong> (Allard 1990) B• Oak - Chestnut - Hickory <strong>Forest</strong> (Ambrose 1990a) BClassification Comments: This forest lacks the dense ericaceous shrub layer typical of other Quercus prinus-dominated forests inthe Blue Ridge escarpment region and commonly has diverse herbaceous composition. It is distinguished from similar forests in theRidge and Valley by lacking Acer saccharum and from Piedmont forests by the lack of Quercus falcata and Quercus stellata, and bythe presence of species more typical of the southern Appalachians (Magnolia fraseri, Halesia tetraptera, and Castanea dentata). Thisassociation was originally defined from the Chattooga Basin Project (S. Simon pers. comm.) and later refined with information fromthe Great Smoky Mountains. The North Carolina Piedmont examples of this association are only montane transition areas, such as theSauratown Mountains and Hanging Rock. It may become more widespread in the Piedmont of Virginia.Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>79
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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ALLIANCE CONCEPTSummary: This allia
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Related Concepts:• Yellow-Poplar:
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virginiana, Sanicula canadensis, De
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Federal Lands: COE (Claiborne Lake)
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ALLIANCE SOURCESReferences: Allard
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Quercus alba and Quercus velutina a
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USFS Ecoregions: 221Hc:CCC, 221He:C
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ALLIANCE CONCEPTSummary: This allia
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- Page 65 and 66: does not occur on wet soils. It occ
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- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
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- Page 119 and 120: USFS Ecoregions: 221Ba:CCC, 221Ha:C
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
- Page 129 and 130: Gettman, R. W. 1974. A floristic su
- Page 131 and 132: Lea, C. 2002a. Vegetation classific
- Page 133 and 134: Pittman, Dr. Albert. Personal commu
- Page 135 and 136: Tobe, J. D., J. E. Fairey, III, and