dominated by deciduous heath species, typically Gaylussacia ursina or Vaccinium stamineum. Other species in the shrub/saplingstratum may include Vaccinium pallidum, Leucothoe recurva, Kalmia latifolia, Castanea dentata, or Acer rubrum var. rubrum. Onrocky sites, Deschampsia flexuosa may be common. This community occurs on exposed upper slopes and ridgetops at elevationsbelow 920 m (3000 feet) in the southern Appalachian Mountains.Environment: This community occurs on exposed upper slopes and ridgetops at elevations below 920 m (3000 feet) in the southernAppalachian Mountains. The presence of Pinus strobus in these forests may be a product of disturbance and subsequent firesuppression. It may have increased its abundance since about 1900.Vegetation: Stands of this forest association typically contain Pinus strobus (contributing 25-75% of the canopy coverage) andQuercus prinus and/or Quercus coccinea (occurring singly or in combination) as 25-75% of the canopy coverage. Open subcanopiesare composed of Oxydendrum arboreum, Acer rubrum var. rubrum, Nyssa sylvatica, and Cornus florida. The shrub stratum isdominated by deciduous heath species, typically Gaylussacia ursina or Vaccinium stamineum. Other species in the shrub/saplingstratum may include Vaccinium pallidum, Leucothoe recurva, Kalmia latifolia, Castanea dentata, or Acer rubrum var. rubrum. Onrocky sites, Deschampsia flexuosa may be common.Dynamics: No informationSimilar Associations:Related Concepts:• Pinus strobus - Quercus coccinea <strong>Forest</strong> (Patterson 1994) ?• Chestnut Oak <strong>Forest</strong> (White Pine Subtype) (Schafale 1998b) ?• Hardwood - White Pine <strong>Forest</strong> (Ambrose 1990a) B• IA6f. Dry White Pine Ridge <strong>Forest</strong> (Allard 1990) B• White Pine - Hardwoods, BR (Pyne 1994) B• White Pine, BR (Pyne 1994) BClassification Comments:CONSERVATION RANKING & RARE SPECIESGRank: G4 (2003-10-23): This community has a restricted range and is uncommon. It is not threatened or particularly vulnerable.Grank changed to G4 on the recommendation of Gary Kauffman, USDA <strong>Forest</strong> Service.High-ranked species: Monotropsis odorata (G3), Thermopsis fraxinifolia (G3?)ELEMENT DISTRIBUTIONRange: This community is known from the escarpment region of the Southern Blue Ridge and may extend into Virginia.Subnations: GA, NC, SC, TN, VA?TNC Ecoregions: 51:C, 52:CUSFS Ecoregions: 221Hb:CCC, 221He:CCC, 222Eo:CCC, M221Db:CCP, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Blue Ridge Parkway?, Carl Sandburg Home, Great Smoky Mountains); USFS (Chattahoochee, Cherokee,Nantahala, Pisgah, <strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, Ambrose 1990a, Patterson 1994, Peet et al. unpubl. data 2002, Pyne 1994, Schafale 1998b, Schafale andWeakley 1990, Southeastern Ecology Working Group n.d.EASTERN WHITE PINE - WHITE OAK - (MOCKERNUT HICKORY) / BEAR HUCKLEBERRY FORESTELEMENT IDENTIFIERSNVC association: Pinus strobus - Quercus alba - (Carya alba) / Gaylussacia ursina <strong>Forest</strong>Database Code: CEGL007517Formation: Mixed needle-leaved evergreen - cold-deciduous forest (I.C.3.N.a)Alliance: Pinus strobus - Quercus (alba, rubra, velutina) <strong>Forest</strong> Alliance (A.401)ELEMENT CONCEPTSummary: This association covers mesic pine-oak-hickory in the Southern Blue Ridge Escarpment and in the Piedmont transition,found below 2900 feet elevation, on protected ridges, mid to upper slopes, and in disturbed bottoms. Canopies are dominated byvariable mixtures of Pinus strobus, Quercus alba, Quercus velutina, Carya alba, and Acer rubrum. Other canopy species may includeLiriodendron tulipifera, Tsuga canadensis, Quercus rubra, Quercus falcata, Quercus prinus, and Magnolia fraseri. Subcanopy andsaplings include canopy species and Cornus florida, Halesia tetraptera, Oxydendrum arboreum, and Nyssa sylvatica. Shrub layers aremoderate to dense, with Gaylussacia ursina and Kalmia latifolia most commonly dominating. Other shrubs include Rhododendronminus, Rhododendron maximum, Symplocos tinctoria, Arundinaria gigantea, Castanea dentata, Sassafras albidum, Amelanchierarborea, Pyrularia pubera, and Hydrangea radiata. The herb stratum is sparse, although ferns (Thelypteris noveboracensis,Dennstaedtia punctilobula and Polystichum acrostichoides) may occasionally dominate. Common herbs include Chimaphilamaculata, Viola hastata, Goodyera pubescens, Maianthemum racemosum, Polygonatum biflorum, Monotropa uniflora, TrilliumVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>76
catesbaei, Desmodium nudiflorum, Eupatorium purpureum, Galium circaezans, Galium latifolium, Galax urceolata, Hexastylisshuttleworthii, Medeola virginiana, Mitchella repens, and Houstonia purpurea.Environment: Stands of this association are found below 2900 feet elevation, on protected ridges, mid to upper slopes, and indisturbed bottoms in the Southern Blue Ridge Escarpment and in the Piedmont transition region. The presence of Pinus strobus inthese forests may be a product of disturbance and subsequent fire suppression. It may have increased its abundance since about 1900.Vegetation: Canopies are dominated by variable mixtures of Pinus strobus, Quercus alba, Carya alba, and Acer rubrum. Othercanopy species may include Liriodendron tulipifera, Tsuga canadensis, Quercus rubra, Quercus prinus, and Magnolia fraseri.Subcanopy and saplings include canopy species and Cornus florida, Halesia tetraptera, Oxydendrum arboreum, and Nyssa sylvatica.Shrub layers are moderate to dense, with Gaylussacia ursina and Kalmia latifolia most commonly dominating. Other shrubs includeRhododendron minus, Rhododendron maximum, Symplocos tinctoria, Arundinaria gigantea, Castanea dentata, Sassafras albidum,Amelanchier arborea, Pyrularia pubera, and Hydrangea radiata. The herb stratum is sparse, although ferns (Thelypterisnoveboracensis, Dennstaedtia punctilobula, and Polystichum acrostichoides) may occasionally dominate. Common herbs includeChimaphila maculata, Viola hastata, Goodyera pubescens, Maianthemum racemosum, Polygonatum biflorum, Monotropa uniflora,Trillium catesbaei, Desmodium nudiflorum, Eupatorium purpureum, Galium circaezans, Galium latifolium, Galax urceolata,Hexastylis shuttleworthii, Medeola virginiana, Mitchella repens, and Houstonia purpurea.Dynamics: No informationSimilar Associations:Related Concepts:• Hardwood - White Pine <strong>Forest</strong> (Ambrose 1990a) B• IA6f. Dry White Pine Ridge <strong>Forest</strong> (Allard 1990) B• Montane Oak-Hickory <strong>Forest</strong> (White Pine Subtype) (Schafale 1998b) ?Classification Comments:CONSERVATION RANKING & RARE SPECIESGRank: G3G4 (2003-10-23): This community is geographically restricted and uncommon within its range. Grank changed to G4 onthe recommendation of Gary Kauffman, USDA <strong>Forest</strong> Service.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This community is known from the escarpment region of the Southern Blue Ridge.Subnations: GA, NC, SC, TN?TNC Ecoregions: 51:CUSFS Ecoregions: M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Great Smoky Mountains); USFS (Chattahoochee, Cherokee, Nantahala, Pisgah, <strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, Ambrose 1990a, <strong>NatureServe</strong> Ecology - Southeastern U.S. unpubl. data, Schafale 1998b, Schafale andWeakley 1990, Southeastern Ecology Working Group n.d.NORTHERN RED OAK - RED MAPLE / SWEET-SHRUB - BUFFALO-NUT / NEW YORK FERN FORESTELEMENT IDENTIFIERSNVC association: Quercus rubra - Acer rubrum / Calycanthus floridus - Pyrularia pubera / Thelypteris noveboracensis <strong>Forest</strong>Database Code: CEGL006192Formation: Lowland or submontane cold-deciduous forest (I.B.2.N.a)Alliance: Quercus alba - (Quercus rubra, Carya spp.) <strong>Forest</strong> Alliance (A.239)ELEMENT CONCEPTSummary: This association includes Quercus rubra forests at intermediate elevations (mostly below 3500 feet, ranging from 2000-4000 feet) in the Southern Blue Ridge Escarpment, and may possibly range into adjacent areas of the Central Appalachians andCumberland Plateau. These forests occur on mostly northern to eastern and southeastern, mid to upper, moderately steep slopes ofintermediate exposure over acidic soils. The canopy is dominated by Quercus rubra, often with Acer rubrum and/or Liriodendrontulipifera codominating, and occasionally with a high component of Quercus alba in the canopy. Other minor canopy species mayinclude Betula lenta, Carya alba, Carya glabra, Halesia tetraptera, Quercus prinus, and Magnolia fraseri. The subcanopy andsapling strata include the canopy species, as well as Halesia tetraptera, Betula lenta, Tsuga canadensis, Cornus florida, Acerpensylvanicum, and Oxydendrum arboreum. The shrub stratum is typically sparse but may have local dominance by Gaylussaciaursina or Rhododendron maximum. Other typical species in the shrub stratum include Castanea dentata, Calycanthus floridus,Pyrularia pubera, Rhododendron calendulaceum, Vaccinium corymbosum, and Viburnum acerifolium. Herbaceous cover is sparse tomoderate but can be species rich. Ferns can be locally dominant, typically Thelypteris noveboracensis and Athyrium filix-femina ssp.asplenioides. Other typical species include Eurybia divaricata (= Aster divaricatus), Carex spp. (e.g., Carex aestivalis, Carex debilis,Carex digitalis, Carex laxiflora var. laxiflora, Carex pensylvanica), Chimaphila maculata (= var. maculata), Desmodium nudiflorum,Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>77
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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• Maritime Oak - Holly Forest / W
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ALLIANCE CONCEPTSummary: This allia
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Related Concepts:• Yellow-Poplar:
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virginiana, Sanicula canadensis, De
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Federal Lands: COE (Claiborne Lake)
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ALLIANCE SOURCESReferences: Allard
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Quercus alba and Quercus velutina a
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USFS Ecoregions: 221Hc:CCC, 221He:C
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- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
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- Page 113 and 114: • Betula nigra - Platanus occiden
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- Page 119 and 120: USFS Ecoregions: 221Ba:CCC, 221Ha:C
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
- Page 129 and 130: Gettman, R. W. 1974. A floristic su
- Page 131 and 132: Lea, C. 2002a. Vegetation classific
- Page 133 and 134: Pittman, Dr. Albert. Personal commu
- Page 135 and 136: Tobe, J. D., J. E. Fairey, III, and