disturbances that fundamentally alter the vegetation structure, floristic composition, and often the physical and chemical structure ofthe soil. Vegetation tends to be dense with a moderately to extremely barren understory. While Pinus echinata is clearly the singlemost dominant tree, other "old-field" Pinus species (e.g., Pinus taeda, Pinus virginiana) and/or other early-successional deciduoustrees (e.g., Acer rubrum, Liquidambar styraciflua, Liriodendron tulipifera) may also be present. Associated woody and herbaceousspecies vary with geography but are typically ruderal or exotic species.Environment: This broadly defined type may develop under a variety of circumstances associated with severe natural and/oranthropogenic disturbance. It is most frequently associated with abandoned agricultural land, unmanaged clearcuts, and burned oreroded areas. These are considered semi-natural forests as they typically result from anthropogenic disturbances which fundamentallyalter the vegetation structure, floristic composition, and often the physical and chemical structure of the soil.Vegetation: Pinus echinata is clearly the single most dominant tree. In addition, other "old-field" Pinus species (e.g., Pinus taeda,Pinus virginiana) and/or other early successional deciduous trees (e.g., Acer rubrum, Liquidambar styraciflua, Liriodendrontulipifera) may also be present. Associated woody and herbaceous species vary with geography but are typically ruderal or exoticspecies.Dynamics: Composition of this community depends more on past disturbance history and adjacent forest composition than on themicro-environment of the site, although this community is limited to flat upland sites and is less common on slopes or protected sites.The community can be found on a variety of upland exposures, but is most well-developed in areas that were heavily farmed and thenabandoned. Over time, the canopy composition shifts as Pinus echinata finishes its short lifespan and other later successional speciesbegin to take advantage of the gaps left by dead and dying pines. At this point, the community can begin to shift to one of a number ofcommunity types that are considered "later successional" than this type and possess a canopy that is dominated by hardwoods ratherthan pines.Similar Associations:• Pinus taeda / Liquidambar styraciflua - Acer rubrum var. rubrum / Vaccinium stamineum <strong>Forest</strong> (CEGL006011) -- occurs insimilar environments with similar disturbance histories, but is dominated by Pinus taeda instead of Pinus echinata.Related Concepts:• IA7a. Xeric Shortleaf Pine - Oak <strong>Forest</strong> (Allard 1990) B• T1A9bI1a. Pinus echinata (Foti et al. 1994) ?Classification Comments: In Kentucky, this vegetation is known only from the eastern part of the state. In Louisiana, thissuccessional vegetation occurs in the Florida parishes and may have a dense shrub understory. In Arkansas, old fields succeed toPinus echinata. Stands have suffered some damage from the Southern Pine Beetle (Dendroctonus frontalis).CONSERVATION RANKING & RARE SPECIESGRank: GNA (ruderal) (2000-4-3): This forest represents a ruderal community resulting from succession following anthropogenicdisturbance of an area. It is not of conservation concern and does not receive a conservation status rank. Stands have suffered somedamage from the Southern Pine Beetle (Dendroctonus frontalis).High-ranked species: No informationELEMENT DISTRIBUTIONRange: This community is found throughout the southeastern United States.Subnations: AL, AR, GA, KY, LA, MO, MS, NC, SC, TN, TX, WV?TNC Ecoregions: 38:C, 39:C, 40:C, 43:C, 44:C, 50:C, 52:P, 53:C, 59:CUSFS Ecoregions: 221J:CC, 222:C, 231A:CC, 231E:CP, 231Ga:CCC, 231Gb:CCC, 231Gc:CCC, 232Bm:CCC, M221A:C?,M221B:C?, M221D:CC, M222A:CCFederal Lands: NPS (Cowpens, Kings Mountain, Little River Canyon?); TVA (Tellico); USFS (Bienville?, Chattahoochee, DanielBoone, Holly Springs?, Mark Twain, Ouachita, Ozark, St. Francis, <strong>Sumter</strong>?, Tombigbee?)ELEMENT SOURCESReferences: Allard 1990, Foti 1994b, Foti et al. 1994, <strong>NatureServe</strong> Ecology - Southeastern U.S. unpubl. data, Southeastern EcologyWorking Group n.d.TULIPTREE - BLACK LOCUST FORESTELEMENT IDENTIFIERSNVC association: Liriodendron tulipifera - Robinia pseudoacacia <strong>Forest</strong>Database Code: CEGL007219Formation: Lowland or submontane cold-deciduous forest (I.B.2.N.a)Alliance: Liriodendron tulipifera <strong>Forest</strong> Alliance (A.236)ELEMENT CONCEPTSummary: This early-successional or semi-natural vegetation occurs in the southern Appalachian Mountains and AppalachianPlateaus. Examples are typical of areas which were once clearcut, old fields, strip-mined, graded for road construction, or otherwisecleared. Stands are typically revegetated from root and stump sprouts and wind dispersed seeds. Canopies are typically dominated byVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 200444Copyright © 2004 <strong>NatureServe</strong>
Liriodendron tulipifera, although other species may also be present. Robinia pseudoacacia is listed as a nominal to indicate theAppalachian distribution of this type. Associated species may vary. Tall shrubs (Rhododendron periclymenoides, Rhododendroncalendulaceum, Kalmia latifolia, Calycanthus floridus) sprout from root stocks and occur as scattered, dense clumps, while shortershrubs (Gaylussacia ursina, Leucothoe fontanesiana, Rubus spp., Vaccinium spp.) can have dense, continuous cover. Composition ofthe herbaceous stratum varies with site conditions and moisture regime and may contain field-adapted species, tolerant of high lightintensities, as well as many shade-tolerant forest herbs. Lycopodium digitatum may also form dense cover.Environment: This community occurs on gentle to moderately steep, middle to upper slopes at approximately 700-1220 m (2300-4000 feet) elevation. Important environmental factors, such as solar irradiation, soil moisture/temperature, and air temperature, varywithin and between sites and are related to the size of the opening, age of the stand, and slope direction (Phillips and Shure 1990).Soils are primarily Hapludults and Dystrochrepts. This successional forest occurs on upland areas in the southern AppalachianMountains and Appalachian Plateaus. It typically occurs as 8- to 16-ha patches in the landscape. These forests are typical of areaswhich were once clearcut, old fields, strip-mined, or cleared by fire or other natural disturbances.Vegetation: Stands of this successional vegetation have canopies which are typically dominated by Liriodendron tulipifera and Acerrubrum, with lesser amounts of Robinia pseudoacacia. Some examples may contain Pinus virginiana. Associated species vary. Therelative absence of Quercus species in these stands indicates more severe disturbance to this type in contrast to Liriodendrontulipifera - Acer rubrum - Quercus spp. <strong>Forest</strong> (CEGL007221). Taller shrubs include Rhododendron periclymenoides, Rhododendroncalendulaceum, Kalmia latifolia, and Calycanthus floridus; shorter shrubs include Gaylussacia ursina, Leucothoe fontanesiana,Rubus spp., and Vaccinium spp. Composition of the herbaceous stratum varies with site conditions and moisture regime and maycontain field-adapted species which are tolerant of high light intensities, as well as many shade-tolerant forest herbs. Lycopodiumdigitatum may also form dense cover.This mainly deciduous forest has a canopy dominated by Liriodendron tulipifera, Acer rubrum, and Robinia pseudoacacia and maycontain lesser amounts of Betula lenta, Cornus florida, Nyssa sylvatica, and Magnolia fraseri. Most of the regeneration is from stumpand root sprouts, however, Liriodendron tulipifera establishment is primarily from seedlings. The upper canopy ranges from 5-9 m,but most of the regeneration is in a shrub/sapling layer at 1-3 m. Cover of woody species may be patchy to dense and is characterizedby clumps of Robinia pseudoacacia and Acer rubrum, occurring as stump sprouts. Scattered thickets of evergreen ericads(Rhododendron and Kalmia) are also typical. Other species occurring as shrubs/saplings include Calycanthus floridus, Halesiacarolina, Pinus strobus, Castanea dentata, Prunus serotina, Pyrularia pubera, Sassafras albidum, Castanea pumila, Hydrangeaarborescens, Viburnum acerifolium, Gaylussacia ursina, Rubus spp., and Vaccinium ssp. Short shrubs (
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
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epeated flooding, this community ma
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USFS Ecoregions: 221Ba:CCC, 221Ha:C
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Classification Comments: On the Ban
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Classification Comments: This veget
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BIBLIOGRAPHYALNHP [Alabama Natural
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DeYoung, H. R. 1979. The white pine
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Gettman, R. W. 1974. A floristic su
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Lea, C. 2002a. Vegetation classific
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Pittman, Dr. Albert. Personal commu
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Tobe, J. D., J. E. Fairey, III, and