Tiarella cordifolia var. collina, Heuchera americana, Stellaria pubera, Podophyllum peltatum, Prenanthes serpentaria, Thalictrumthalictroides, Chrysogonum virginianum var. virginianum, Hepatica nobilis var. obtusa, Thelypteris noveboracensis, and Botrychiumvirginianum. Exact composition varies locally with position on slope and nature of soil. Western Piedmont sites often have increasingimportance of Tsuga canadensis, Rhododendron spp., and other species that are more typical of the Southern Blue Ridge.Environment: Examples of this association predominantly occur on steep but sheltered slopes adjacent to creeks or rivers in thePiedmont. They can occur further upslope, but occurrences are much more likely as one gets closer to streams.Vegetation: The canopy of stands of this association is dominated by mesophytic trees such as Fagus grandifolia, Quercus rubra,Liriodendron tulipifera, Acer rubrum, and in the western Piedmont, Tsuga canadensis. Typical understory trees include Cornusflorida, Oxydendrum arboreum, Acer rubrum, and Ilex opaca. Shrub species may include Vaccinium stamineum, Viburnumrafinesquianum, Euonymus americana, and sometimes Kalmia latifolia. The herb layer is often moderately dense and diverse, thoughit may be sparse under heavy shade. Herb species may include Polystichum acrostichoides, Viola spp., Dichanthelium spp. (=Panicum spp.), Galium circaezans, Hexastylis arifolia, Hexastylis minor, Desmodium nudiflorum, Erythronium umbilicatum ssp.umbilicatum, Chamaelirium luteum, Epifagus virginiana, Tiarella cordifolia var. collina, Heuchera americana, Stellaria pubera,Podophyllum peltatum, Prenanthes serpentaria, Thalictrum thalictroides, Chrysogonum virginianum var. virginianum, Hepaticanobilis var. obtusa, Thelypteris noveboracensis, and Botrychium virginianum (Schafale and Weakley 1990). Exact composition varieslocally with position on slope and nature of soil. Western Piedmont sites often have increasing importance of Tsuga canadensis,Rhododendron spp., and other species that are more typical of the Southern Blue Ridge.Dynamics: Under natural conditions these forests are uneven-aged, with old trees present. Reproduction occurs primarily in canopygaps. Rare, severe natural disturbances such as wind storms may allow pulses of increased regeneration and allow the less shadetolerantspecies to remain in the community. However, Skeen, Carter, and Ragsdale (1980) argued that even the shade-intolerantLiriodendron could reproduce enough in gaps to persist in the climax Piedmont forests (Schafale and Weakley 1990).The natural fire regime of the Piedmont is not known but fires certainly occurred periodically. Because Mesic Mixed Hardwood<strong>Forest</strong>s generally occur in moist and topographically sheltered sites, they probably burned only rarely and with low intensity (Schafaleand Weakley 1990).Disturbed areas have increased amounts of pines and weedy hardwoods such as Liriodendron tulipifera and Liquidambar styraciflua.Many areas have been selectively cut many times and have increased importance of Fagus grandifolia and other noncommercialhardwoods relative to oaks (Schafale and Weakley 1990). Other areas that were disturbed in the distant past may be younger and,therefore, may have a higher proportion of oaks with beeches mainly in the understory.Similar Associations:• Fagus grandifolia - Quercus rubra / Ostrya virginiana - Acer (barbatum, leucoderme) / Actaea racemosa - Sanguinariacanadensis <strong>Forest</strong> (CEGL008466) -- the basic Piedmont equivalent.Related Concepts:• Fagus grandifolia - Quercus (alba, rubra) - Liriodendron tulipifera / Ilex opaca var. opaca - (Asimina triloba) <strong>Forest</strong> (Flemingpers. comm.) ?• Fagus grandifolia - Quercus rubra - Quercus alba / Carpinus caroliniana <strong>Forest</strong> (Lea 2002a) ?Classification Comments:CONSERVATION RANKING & RARE SPECIESGRank: G3G4 (2001-1-18): While not as extensive as the oak-hickory forests, Mesic Mixed Hardwood <strong>Forest</strong> communities arefairly common. Their occurrence on steep sites has allowed many of them to escape, until recently, with less disturbance than mostupland communities (Schafale and Weakley 1990). Some examples with old forest can be found. Selective timbering may havedecreased the importance value of more desirable hardwoods (e.g., Quercus rubra). Some examples are protected in the BirkheadMountain Wilderness Area and other parts of the Uwharrie <strong>National</strong> <strong>Forest</strong>.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This association is found in the Piedmont of the southeastern United States, from Maryland to Georgia.Subnations: GA, MD, NC, SC, VATNC Ecoregions: 52:C, 58:CUSFS Ecoregions: 231Aa:CCC, 231Ae:CCCFederal Lands: DOD (Falls Lake, Jordan Lake, Kerr Reservoir); NPS (Guilford Courthouse, Ninety Six, Thomas Stone); USFS(Oconee?, <strong>Sumter</strong>?, Uwharrie)ELEMENT SOURCESReferences: Fleming 2001, Fleming et al. 2001, Fleming pers. comm., LeGrand and Dalton 1987, Lea 2002a, Nehmeth 1968,Oosting 1942, Peet and Christensen 1980, Peet et al. unpubl. data 2002, Schafale and Weakley 1990, Skeen et al. 1980, SoutheasternEcology Working Group n.d.Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>66
WHITE OAK - MOCKERNUT HICKORY / AMERICAN STRAWBERRY-BUSH / ARROWLEAF HEARTLEAF FORESTELEMENT IDENTIFIERSNVC association: Quercus alba - Carya alba / Euonymus americana / Hexastylis arifolia <strong>Forest</strong>Database Code: CEGL006227Formation: Lowland or submontane cold-deciduous forest (I.B.2.N.a)Alliance: Quercus alba - (Quercus rubra, Carya spp.) <strong>Forest</strong> Alliance (A.239)ELEMENT CONCEPTSummary: This type represents mesic, subacid oak-hickory forests of the southern Piedmont and possibly adjacent ecoregions.Stands of this association typically exhibit heavy dominance by Quercus alba and Carya alba. The canopy or subcanopy may alsocontain Fagus grandifolia, Liquidambar styraciflua, Liriodendron tulipifera, Carya ovalis, Prunus serotina, Acer rubrum, Quercusnigra, Quercus falcata, Quercus velutina, Oxydendrum arboreum, Fraxinus spp., Halesia sp., Nyssa sylvatica, and Cornus florida.Some stands may contain a minor component of Pinus echinata. Shrubs include Euonymus americana, Rhododendron canescens,Calycanthus floridus, Aesculus pavia, Hydrangea quercifolia, Vaccinium elliottii, Vaccinium pallidum, Vaccinium stamineum,Asimina parviflora, and Asimina parviflora. Vines such as Toxicodendron radicans, Smilax rotundifolia, Smilax glauca,Parthenocissus quinquefolia, and Vitis rotundifolia are abundant. Some typical herbs are Hexastylis arifolia, Mitchella repens,Polystichum acrostichoides, Dioscorea villosa, Elephantopus tomentosus, Maianthemum racemosum, Coreopsis major, Desmodiumnudiflorum, and Hypoxis hirsuta.Environment: This community occupies protected low slopes in the acidic Piedmont where creeks have cut small draws or valleys.Vegetation: Stands of this association typically exhibit heavy dominance by Quercus alba and Carya alba. The canopy orsubcanopy may also contain Fagus grandifolia, Liquidambar styraciflua, Liriodendron tulipifera, Carya ovalis, Prunus serotina, Acerrubrum, Quercus nigra, Quercus falcata, Quercus velutina, Oxydendrum arboreum, Fraxinus spp., Halesia sp., Nyssa sylvatica, andCornus florida. Some stands may contain a minor component of Pinus echinata. Shrubs include Euonymus americana, Rhododendroncanescens, Calycanthus floridus, Aesculus pavia, Hydrangea quercifolia, Vaccinium elliottii, Vaccinium pallidum, Vacciniumstamineum, Asimina parviflora, and Asimina parviflora. Vines such as Toxicodendron radicans, Smilax rotundifolia, Smilax glauca,Parthenocissus quinquefolia, and Vitis rotundifolia are abundant. Some typical herbs are Hexastylis arifolia, Mitchella repens,Polystichum acrostichoides, Dioscorea villosa, Elephantopus tomentosus, Maianthemum racemosum, Coreopsis major, Desmodiumnudiflorum, and Hypoxis hirsuta (Golden 1979).Dynamics: This community typically exists on sheltered gentle slopes leading down to drainageways in the Piedmont. It is mostlikely not fire-tolerant.Similar Associations:• Quercus alba - Quercus (rubra, coccinea) - Carya (alba, glabra) / Vaccinium pallidum Piedmont Dry-Mesic <strong>Forest</strong>(CEGL008475) -- similar with a more northerly range.• Quercus alba - Quercus rubra - Carya glabra - Carya ovata / Viburnum rafinesquianum / Viola tripartita <strong>Forest</strong> (CEGL007236)-- relationship is unclear.• Quercus falcata - Quercus alba - Carya alba / Oxydendrum arboreum / Vaccinium stamineum <strong>Forest</strong> (CEGL007244) -- a lessmesic type with range overlap in the southern Piedmont.• Quercus rubra - Quercus alba - Carya glabra / Geranium maculatum <strong>Forest</strong> (CEGL007237)Related Concepts:• White Oak (Golden 1979) ?Classification Comments: Based on data in Golden (1979; 6 stands sampled). This association needs clarification and expansion,and subsequent range assessment. In particular, compare to Quercus rubra - Quercus alba - Carya glabra / Geranium maculatum<strong>Forest</strong> (CEGL007237) and Quercus alba - Quercus (rubra, coccinea) - Carya (alba, glabra) / Vaccinium pallidum Piedmont Dry-Mesic <strong>Forest</strong> (CEGL008475). Golden's data may actually represent a variety of Quercus alba-dominated associations. Moreinformation is needed.CONSERVATION RANKING & RARE SPECIESGRank: G4G5 (2002-9-3): Global rank changed from G5? to G4G5 to better reflect its rarity and the possible uncertainty. This is asomewhat widespread type, but is not secure across its range. The Piedmont is an area of rapid population growth, and at a minimumthis forest is threatened by removal of commercially valuable timber species (e.g., Quercus alba), conversion to other forest types,and fragmentation and removal in areas of human population growth.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This association is found in the Piedmont from South Carolina to Alabama.Subnations: AL, GA, SCTNC Ecoregions: 52:CUSFS Ecoregions: 231A:CC, 231D:CCFederal Lands: NPS (Cowpens); USFS (Oconee, <strong>Sumter</strong>?)Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>67
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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• Maritime Oak - Holly Forest / W
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ALLIANCE CONCEPTSummary: This allia
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Related Concepts:• Yellow-Poplar:
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- Page 39 and 40: Federal Lands: COE (Claiborne Lake)
- Page 41 and 42: ALLIANCE SOURCESReferences: Allard
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- Page 45 and 46: USFS Ecoregions: 221Hc:CCC, 221He:C
- Page 47 and 48: ALLIANCE CONCEPTSummary: This allia
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- Page 53 and 54: the associated species. More Appala
- Page 55 and 56: ALLIANCE DISTRIBUTIONRange: This al
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- Page 59 and 60: elevation environments such as Leio
- Page 61 and 62: CULTIVATED FORESTEASTERN WHITE PINE
- Page 63 and 64: Federal Lands: DOD (Arnold, Fort Be
- Page 65 and 66: does not occur on wet soils. It occ
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- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
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- Page 101 and 102: ELEMENT SOURCESReferences: Allard 1
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- Page 109 and 110: ELEMENT DISTRIBUTIONRange: This for
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- Page 113 and 114: • Betula nigra - Platanus occiden
- Page 115 and 116: Formation: Temporarily flooded temp
- Page 117 and 118: epeated flooding, this community ma
- Page 119 and 120: USFS Ecoregions: 221Ba:CCC, 221Ha:C
- Page 121 and 122: Classification Comments: On the Ban
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
- Page 129 and 130: Gettman, R. W. 1974. A floristic su
- Page 131 and 132: Lea, C. 2002a. Vegetation classific
- Page 133 and 134: Pittman, Dr. Albert. Personal commu
- Page 135 and 136: Tobe, J. D., J. E. Fairey, III, and