• White Pine - Chestnut Oak: 51 (Eyre 1980) ?• White Pine - Hardwoods, BR (Pyne 1994) ?• White Pine <strong>Forest</strong> (Schafale and Weakley 1990) I• White Pine, BR (Pyne 1994) ?Classification Comments: Isolated stands with Pinus strobus are found in the vicinity of Clifty, Kentucky. These occur onsandstones of the Dripping Springs escarpment (Logan, Muhlenburg, Todd counties) and are presumably more xeric. Their placementis unclear.ALLIANCE DISTRIBUTIONRange: This alliance is found in Georgia, North Carolina, South Carolina, Tennessee, and may also be found in Virginia (?).Subnations: GA, MD?, NC, SC, TN, VA, WVTNC Ecoregions: 50:C, 51:C, 52:C, 59:CUSFS Ecoregions: 221Hb:CCC, 221He:CCC, 222Eo:CCC, 231Aa:CC?, 231Ae:CCC, 231Ak:CCC, 231Al:CC?, 231Ap:CCC,M221Aa:CCC, M221Ab:CCC, M221Da:CCC, M221Db:CCP, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Blue Ridge Parkway?, Carl Sandburg Home, Great Smoky Mountains); USFS (Chattahoochee, Cherokee?,George Washington, Jefferson, Nantahala, Pisgah, <strong>Sumter</strong>)ALLIANCE SOURCESReferences: Allard 1990, Ambrose 1990a, DuMond 1970, Eyre 1980, Gattis 1992, Patterson 1994, Pyne 1994, Schafale andWeakley 1990I.C.3.N.A. PINUS STROBUS - QUERCUS (ALBA, RUBRA, VELUTINA) FOREST ALLIANCE (A.401)EASTERN WHITE PINE - (WHITE OAK, NORTHERN RED OAK, BLACK OAK) FORESTALLIANCEALLIANCE CONCEPTSummary: This alliance occurs from the western Great Lakes to the northeastern United States and south to the southernAppalachian Mountains. The overstory is a mix of evergreen and deciduous trees which form a moderately closed to closed canopy.Pinus strobus is a consistent constituent of the canopy and usually occurs as supercanopy trees, as well. Quercus alba, Quercus rubra,and Quercus velutina are also important canopy trees along with minor amounts of Acer rubrum, Carya alba, Liriodendron tulipifera,Pinus resinosa, Pinus banksiana (in Wisconsin), Populus tremuloides (in the northern parts of this alliance's range), Quercusellipsoidalis (in the northwest), and Tsuga canadensis, and Quercus prinus (in the southeast). Subcanopy trees can include Carpinuscaroliniana, Cornus florida, Hamamelis virginiana, Halesia tetraptera, Oxydendrum arboreum, and Nyssa sylvatica. The shrub layeris often well-developed with Gaylussacia spp., Kalmia latifolia, Rubus spp., and Vaccinium spp. most commonly dominant. Othershrubs can include Corylus americana, Gaultheria procumbens, Rhododendron maximum, and Sassafras albidum, and in the Ridgeand Valley, Viburnum rafinesquianum and Viburnum prunifolium. The herb stratum is sparse to moderate, but can be quite speciesrich, especially in the Southern Blue Ridge and Ridge and Valley, where typical species include Ageratina altissima, Amphicarpaeabracteata, Brachyelytrum erectum, Carex communis, Carex platyphylla, Carex woodii, Carex pensylvanica, Chimaphila maculata,Desmodium nudiflorum, Eupatorium purpureum, Galax urceolata, Galium latifolium, Galium circaezans, Geranium maculatum,Goodyera pubescens, Hexastylis shuttleworthii, Hieracium venosum, Houstonia purpurea, Maianthemum racemosum, Maianthemumcanadense, Medeola virginiana, Mitchella repens, Monotropa uniflora, Poa cuspidata, Polygonatum biflorum, Polystichumacrostichoides, Trillium catesbaei, and Viola hastata. Stands of this alliance are dry-mesic to mesic forests found on acidic, relativelynutrient-poor, sandy loam to sandy soil on a variety of topographic positions. In the upper Midwest, most stands are on flat to rollingtopography on outwash plains or moraines. In the Southern Blue Ridge, they occur on mid to lower slopes in the lower elevations(below 3000 feet) on protected ridges, and in disturbed bottoms. In the Ridge and Valley of Virginia, these forests are found onprotected ravines, with rocky soils developed over shale, sandstone, or other sedimentary rock.Related Concepts:• Pinus strobus - Pinus resinosa forest (No. 36) (Vankat 1990) I• Coastal <strong>Forest</strong>/Woodland (Swain and Kearsley 2001) ?• Dry - Mesic Acidic Central <strong>Forest</strong> (Smith 1991) ?• Dry white pine (hemlock) - oak forest (Fike 1999) ?• Dry-Mesic Oak--Hickory <strong>Forest</strong> (Schafale and Weakley 1990) I• Eastern White Pine: 21 (Eyre 1980) I• Hardwood - White Pine <strong>Forest</strong> (Ambrose 1990a) I• Northern Dry-mesic <strong>Forest</strong> (Curtis 1959) I• White Pine - Chestnut Oak: 51 (Eyre 1980) I• White Pine - Northern Red Oak - Red Maple: 20 (Eyre 1980) ?Classification Comments: In the Appalachians, these forests are typically transitional between the more mesic, protected coveforests and the more xeric, exposed pine - oak forests with Quercus prinus and Quercus coccinea. Stands of this alliance are midsuccessionalbut long-lasting and require repeated disturbances to regenerate (MNNHP 1993). Isolated stands of Pinus strobus withVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>20
Quercus alba and Quercus velutina and scattered Fagus grandifolia over Kalmia latifolia are found on steep slopes of the WesternHighland Rim of (Cheatham and Dickson counties; Chester 1980). Similar isolated stands with Pinus strobus are found in the vicinityof Clifty, Kentucky. Their status is unclear as well. These occur on sandstones of the Dripping Springs escarpment (Logan,Muhlenburg, Todd counties). "These are along Clifty Creek Gorge and Rocky Creek/Lake Malone State Park area; no botanicalperson has reported from here for a long time. The environment is not necessarily more xeric." (Julian Campbell, TNC-KYFO).ALLIANCE DISTRIBUTIONRange: This alliance is found in northern Minnesota, Wisconsin, Michigan, northern Illinois, Indiana, Connecticut, Maine,Maryland, Massachusetts, New Jersey, New York, Pennsylvania, Rhode Island, Virginia, West Virginia, northern Georgia, westernNorth Carolina, western South Carolina, and eastern Tennessee (?). It is also found in Canada.Subnations: GA, IL, IN, MA, MD, ME, MI, MN, NC, NH, NJ?, NY, ON, PA, QC?, RI, SC, TN?, VA?, VT, WI, WVTNC Ecoregions: 44:P, 46:C, 47:C, 48:C, 49:C, 50:P, 51:C, 52:P, 59:C, 60:C, 61:C, 62:C, 63:C, 64:CUSFS Ecoregions: 212Aa:CPP, 212Ab:CPP, 212Ba:CPP, 212Bb:CPP, 212Ca:CCP, 212Cb:CCC, 212Da:CCC, 212Db:CCC,212Dc:CCC, 212Eb:CCC, 212Ec:CCP, 212Ed:CCP, 212Fa:CCP, 212Fb:CCC, 212Fc:CCC, 212Fd:CCC, 212Ga:CCP, 212Gb:CCP,212Hb:CCP, 212He:CCC, 212Hi:CCP, 212Hj:CCC, 212Hl:CCC, 212Hm:CCP, 212Ho:CCC, 212Hp:CCC, 212Hq:CCC, 212Hr:CCP,212Hs:CCP, 212Ht:CCP, 212Hu:CCC, 212Hv:CCC, 212Hw:CCC, 212Hx:CCP, 212Hy:CCC, 212Ja:CCP, 212Jb:CCC, 212Jc:CCP,212Jf:CCP, 212Jj:CCP, 212Jl:CCP, 212Jm:CCC, 212Jn:CCP, 212Jr:CCC, 212Ka:CCP, 212Kb:CCC, 212La:CPP, 212Lb:CP?,212Mb:CPP, 212Na:CPP, 212Nb:CPP, 212Nc:CPP, 212Oa:CCC, 212Oc:CCC, 221Aa:CCC, 221Ab:CCP, 221Ac:CCC, 221Ae:CCC,221Af:CCP, 221Ag:CCC, 221Ah:CCC, 221Ai:CCC, 221Aj:CCP, 221Ak:CCC, 221Al:CCC, 221Am:CCP, 221Ba:CCC, 221Bb:CCP,221Bc:CCC, 221Bd:CCC, 221Db:CPP, 221Ea:CCC, 221Ec:CCC, 221Fa:CCC, 222Eg:CCC, 222Ic:CCP, 222Id:CCP, 222If:CCP,222Ja:CCC, 222Jc:CCC, 222Jd:CCC, 222Je:CCC, 222Jg:CCC, 222Jj:CCC, 222Ka:CCC, 222Kh:CCC, 222Lc:CCC, 222Lf:CCC,222Mc:CCC, 222Md:CCC, 222Me:CCC, 232Aa:CCC, M212Ac:CC?, M212Ad:CCC, M212Ae:CCP, M212Ag:CCC, M212Ba:CCP,M212Bb:CCC, M212Bc:CCC, M212Bd:CCC, M212Ca:CCP, M212Cb:CCC, M212Cc:CCC, M212Cd:CCP, M212Db:CCP,M212Dc:CCC, M212De:CCC, M212Ea:CCC, M212Eb:CCC, M221Aa:CCC, M221Ab:CCC, M221Ac:CCC, M221Ad:CCC,M221Ba:CC?, M221Bb:CCC, M221Bc:CC?, M221Bd:CCC, M221Be:CCC, M221Bf:CCC, M221Ca:CC?, M221Cb:CCP,M221Da:CCC, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Acadia, Blue Ridge Parkway?, Great Smoky Mountains?); USFS (Chattahoochee, Cherokee?, GeorgeWashington, Nantahala, Pisgah, <strong>Sumter</strong>)ALLIANCE SOURCESReferences: Ambrose 1990a, Chester and Scott 1980, Curtis 1959, Eyre 1980, Faber-Langendoen et al. 1996, Fike 1999, MNNHP1993, Rawinski et al. 1996, Schafale and Weakley 1990, Smith 1991, Swain and Kearsley 2001, Vankat 1990I.C.3.N.A. PINUS TAEDA - QUERCUS (ALBA, FALCATA, STELLATA) FOREST ALLIANCE (A.404)LOBLOLLY PINE - (WHITE OAK, SOUTHERN RED OAK, POST OAK) FOREST ALLIANCEALLIANCE CONCEPTSummary: This alliance encompasses loblolly pine - oak forests of the Coastal Plain and some adjacent provinces of the easternUnited States. The canopy is dominated by Pinus taeda with some combination of the nominal oaks. More mesic examples tend to becodominated by Quercus alba, while dry to dry-mesic examples are usually codominated by Quercus falcata. Associated species varyby geography, substrate, and exposure. Described members of this alliance are found sporadically, ranging from the North AtlanticCoast of Delaware, through the Chesapeake Lowlands of Maryland and Virginia to the West Gulf and Upper West Gulf coastal plainsof eastern Texas and Arkansas where they are most common. These forests are apparently absent from the Mid-Atlantic Coastal Plainof North and South Carolina, but are documented in the South Atlantic Coastal Plain of Georgia. Successional and/or semi-naturalexamples are known from the East Gulf and Upper East Gulf coastal plains. Within the longleaf pine belt, these forests can occurnaturally on fire-protected areas such as topographically isolated hilltops, mid to lower slopes, protected ravines, broad flats andsecond bottoms. In some cases, they are successional forests on broad uplands following clearing or alteration of natural forests,especially those historically dominated by Pinus palustris A broad range of associated species may be present in this type, includingCarya alba, Carya texana, Nyssa sylvatica, Liquidambar styraciflua, Carya cordiformis, Magnolia grandiflora, Fagus grandifolia,Quercus velutina, Quercus michauxii, Quercus pagoda, and Acer rubrum. The subcanopy can include canopy species, as well as Ilexopaca var. opaca, Ostrya virginiana, Carpinus caroliniana, Cornus florida, and others. Callicarpa americana, Symplocos tinctoria,Morella cerifera (= Myrica cerifera), Vaccinium elliottii, Viburnum dentatum, and Viburnum acerifolium are common shrub species.Herbaceous species that may be present include Polystichum acrostichoides, Athyrium filix-femina ssp. asplenioides, Phegopterishexagonoptera, Prenanthes altissima, Spigelia marilandica, Mitchella repens, Podophyllum peltatum, Phlox divaricata, Tipulariadiscolor, Arisaema triphyllum, Erigeron pulchellus, Lilium michauxii, Chasmanthium laxum, Chasmanthium sessiliflorum, andMelica mutica.Related Concepts:• Calcareous <strong>Forest</strong> (Smith 1996a) I• Dry-Mesic Mixed Oak - Pine <strong>Forest</strong> (Wieland 1994b) ?• IA6e. Loblolly Pine - Shortleaf Pine - Oak <strong>Forest</strong> (Allard 1990) IVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>21
- Page 3: 1 NatureServe is an international o
- Page 11 and 12: types. However, when necessary, mod
- Page 13 and 14: [Association name = floristic nomin
- Page 15 and 16: Generally 5 or fewer occurrences an
- Page 17 and 18: All scientific names for vascular s
- Page 19 and 20: Summary: A short description of the
- Page 21 and 22: Loucks, O. 1996. 100 years after Co
- Page 23 and 24: ALLIANCES BY US NATIONAL VEGETATION
- Page 25 and 26: virginiana, and Acer pensylvanicum
- Page 27 and 28: virginiana, Juniperus virginiana va
- Page 29 and 30: • Appalachian pine-oak forest (Ev
- Page 31 and 32: • Maritime Oak - Holly Forest / W
- Page 33 and 34: ALLIANCE CONCEPTSummary: This allia
- Page 35 and 36: Related Concepts:• Yellow-Poplar:
- Page 37 and 38: virginiana, Sanicula canadensis, De
- Page 39 and 40: Federal Lands: COE (Claiborne Lake)
- Page 41: ALLIANCE SOURCESReferences: Allard
- Page 45 and 46: USFS Ecoregions: 221Hc:CCC, 221He:C
- Page 47 and 48: ALLIANCE CONCEPTSummary: This allia
- Page 49 and 50: USFS Ecoregions: 221Eb:PP?, M221Ab:
- Page 51 and 52: Federal Lands: NPS (Great Smoky Mou
- Page 53 and 54: the associated species. More Appala
- Page 55 and 56: ALLIANCE DISTRIBUTIONRange: This al
- Page 57 and 58: USFS Ecoregions: 221Ha:CCC, 221Hc:C
- Page 59 and 60: elevation environments such as Leio
- Page 61 and 62: CULTIVATED FORESTEASTERN WHITE PINE
- Page 63 and 64: Federal Lands: DOD (Arnold, Fort Be
- Page 65 and 66: does not occur on wet soils. It occ
- Page 67 and 68: Liriodendron tulipifera, although o
- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
- Page 73 and 74: Similar Associations:• Pinus echi
- Page 75 and 76: • Pinus echinata - Quercus stella
- Page 77 and 78: this association occurs in edaphica
- Page 79 and 80: Vegetation: This forest typically h
- Page 81 and 82: strobus can have high coverage and
- Page 83 and 84: CONSERVATION RANKING & RARE SPECIES
- Page 85 and 86: • Quercus alba - Carya alba / Euo
- Page 87 and 88: muehlenbergii. In addition, Acer ba
- Page 89 and 90: WHITE OAK - MOCKERNUT HICKORY / AME
- Page 91 and 92: CONSERVATION RANKING & RARE SPECIES
- Page 93 and 94:
Vegetation: This association encomp
- Page 95 and 96:
ELEMENT DISTRIBUTIONRange: This com
- Page 97 and 98:
• Quercus prinus - Carya spp. - Q
- Page 99 and 100:
catesbaei, Desmodium nudiflorum, Eu
- Page 101 and 102:
ELEMENT SOURCESReferences: Allard 1
- Page 103 and 104:
High-ranked species: No information
- Page 105 and 106:
Dynamics: No informationSimilar Ass
- Page 107 and 108:
protrusa, Grammitis nimbata (= Micr
- Page 109 and 110:
ELEMENT DISTRIBUTIONRange: This for
- Page 111 and 112:
COMMON WATER-WILLOW HERBACEOUS VEGE
- Page 113 and 114:
• Betula nigra - Platanus occiden
- Page 115 and 116:
Formation: Temporarily flooded temp
- Page 117 and 118:
epeated flooding, this community ma
- Page 119 and 120:
USFS Ecoregions: 221Ba:CCC, 221Ha:C
- Page 121 and 122:
Classification Comments: On the Ban
- Page 123 and 124:
Classification Comments: This veget
- Page 125 and 126:
BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128:
DeYoung, H. R. 1979. The white pine
- Page 129 and 130:
Gettman, R. W. 1974. A floristic su
- Page 131 and 132:
Lea, C. 2002a. Vegetation classific
- Page 133 and 134:
Pittman, Dr. Albert. Personal commu
- Page 135 and 136:
Tobe, J. D., J. E. Fairey, III, and