• Pinus echinata - Quercus alba / Vaccinium pallidum / Hexastylis arifolia - Chimaphila maculata <strong>Forest</strong> (CEGL008427) -- arelated mixed type.• Quercus alba - Carya alba / Euonymus americana / Hexastylis arifolia <strong>Forest</strong> (CEGL006227) -- a more mesic type with rangeoverlap in the southern Piedmont.• Quercus alba - Quercus falcata / Vaccinium (arboreum, hirsutum, pallidum) <strong>Forest</strong> (CEGL008567) -- more eastern and montanein distribution.• Quercus falcata - Quercus alba - Quercus stellata - Quercus velutina <strong>Forest</strong> (CEGL005018)Related Concepts:• Black Oak: 110 (Eyre 1980) B• IA6i. Interior Upland Dry-Mesic Oak - Hickory <strong>Forest</strong> (Allard 1990) B• Mesotrophic <strong>Forest</strong> (Rawinski 1992) ?• Southern Red Oak, HR (Pyne 1994) B• Submesic Broadleaf Deciduous <strong>Forest</strong> (Ambrose 1990a) B• White Oak - Black Oak - Northern Red Oak: 52 (Eyre 1980) B• White Oak - Mixed Hardwoods, HR (Pyne 1994) B White Oak - Mixed Oak - Hickory, HR (Pyne 1994) B• White Oak: 53 (Eyre 1980) BClassification Comments: The limits of the range of this type needs to be clarified in Kentucky.CONSERVATION RANKING & RARE SPECIESGRank: G4G5 (2002-10-15): This is not a rare forest type, although most examples have been impacted by removal of the morevaluable timber species (e.g., Quercus alba), and remaining ones on private land are highly vulnerable to canopy removal andconversion to other forest types or other land uses.High-ranked species: Amorpha schwerinii (G3G4), Brickellia cordifolia (G2G3), Monotropsis odorata (G3), Rhus michauxii (G2)ELEMENT DISTRIBUTIONRange: This southern red oak - white oak dry forest is found in the Piedmont of Georgia, South Carolina, North Carolina, andVirginia, and in the interior uplands and Cumberland Plateau of Kentucky and Tennessee. It has also been reported from the UpperEast Gulf Coastal Plain of Mississippi, Tennessee and Georgia.Subnations: AL, GA, KY, MS, NC, SC, TN, VA?TNC Ecoregions: 43:C, 44:C, 50:C, 52:C, 53:?USFS Ecoregions: 221Hc:CCC, 222Cg:CCC, 222Eb:CCC, 231Ae:CCCFederal Lands: DOD (Arnold, Fort Benning, Fort Gordon?); DOE (Oak Ridge); NPS (Cowpens, Guilford Courthouse, KingsMountain, Little River Canyon?, Ninety Six, Shiloh); USFS (Daniel Boone, Holly Springs?, Oconee, <strong>Sumter</strong>, Talladega, Uwharrie);USFWS (Eufaula)ELEMENT SOURCESReferences: ALNHP 2002, Allard 1990, Ambrose 1990a, Evans 1991, Eyre 1980, Golden 1979, <strong>NatureServe</strong> Ecology -Southeastern U.S. unpubl. data, Oberholster 1993, Oosting 1942, Peet and Christensen 1980, Peet et al. unpubl. data 2002, Pyne1994, Rawinski 1992, Schafale and Weakley 1990, Southeastern Ecology Working Group n.d.WHITE OAK - NORTHERN RED OAK - PIGNUT HICKORY - SHAGBARK HICKORY / DOWNY ARROW-WOOD / THREE-PARTED YELLOW VIOLET FORESTELEMENT IDENTIFIERSNVC association: Quercus alba - Quercus rubra - Carya glabra - Carya ovata / Viburnum rafinesquianum / Viola tripartita <strong>Forest</strong>Database Code: CEGL007236Formation: Lowland or submontane cold-deciduous forest (I.B.2.N.a)Alliance: Quercus alba - (Quercus rubra, Carya spp.) <strong>Forest</strong> Alliance (A.239)ELEMENT CONCEPTSummary: This association accommodates basic, dry-mesic oak-hickory forests of the Piedmont. Stands would be expected tocontain Quercus alba, Quercus rubra, Carya glabra, and Carya ovata in the canopy. Other species indicative of mafic orcircumneutral conditions would also be present, such as Viburnum rafinesquianum, Cercis canadensis, Quercus shumardii, andpossibly other species.Environment: No informationVegetation: Stands of this type would be expected to contain Quercus alba, Quercus rubra, Carya glabra, and Carya ovata in thecanopy, Viburnum rafinesquianum as a typical shrub, and Viola tripartita as a typical herb. More information is needed.Dynamics: No informationSimilar Associations:Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>62
• Quercus alba - Carya alba / Euonymus americana / Hexastylis arifolia <strong>Forest</strong> (CEGL006227) -- has an overlapping range, but ispresumably acidic.• Quercus alba - Carya ovata / Cercis canadensis <strong>Forest</strong> (CEGL007232)• Quercus alba - Quercus (rubra, coccinea) - Carya (alba, glabra) / Vaccinium pallidum Piedmont Dry-Mesic <strong>Forest</strong>(CEGL008475) -- is a related non-mafic type.• Quercus alba - Quercus stellata - (Quercus rubra) - Carya carolinae-septentrionalis / Acer leucoderme - Cercis canadensis<strong>Forest</strong> (CEGL007773) -- is related mafic type with overlapping distribution, apparently somewhat drier.Related Concepts: No informationClassification Comments: After Golden (1979). [MP 1999-02 note: this type, based on its nominals, does NOT correspond to aGolden type. Viburnum rafinesquianum and Viola tripartita are not present in Golden's data. More information is needed. Golden's"White Oak" type is explicitly represented by CEGL006227.] This association needs clarification and expansion, and subsequentrange assessment. Compare to Quercus alba - Quercus (rubra, coccinea) - Carya (alba, glabra) / Vaccinium pallidum Piedmont Dry-Mesic <strong>Forest</strong> (CEGL008475) and to Quercus alba - Carya ovata / Cercis canadensis <strong>Forest</strong> (CEGL007232).CONSERVATION RANKING & RARE SPECIESGRank: G3G4 (1997-8-14): No informationHigh-ranked species: No informationELEMENT DISTRIBUTIONRange: This association could be found in the Piedmont ecoregion from Alabama to Virginia.Subnations: AL, GA, NC, SC, VA?TNC Ecoregions: 52:CUSFS Ecoregions: 231:CFederal Lands: USFS (Oconee?, <strong>Sumter</strong>?, Uwharrie)ELEMENT SOURCESReferences: Golden 1979, Peet and Christensen 1980, Peet et al. unpubl. data 2002, Schafale and Weakley 1990, SoutheasternEcology Working Group n.d.WHITE OAK - SHAGBARK HICKORY / REDBUD FORESTELEMENT IDENTIFIERSNVC association: Quercus alba - Carya ovata / Cercis canadensis <strong>Forest</strong>Database Code: CEGL007232Formation: Lowland or submontane cold-deciduous forest (I.B.2.N.a)Alliance: Quercus alba - (Quercus rubra, Carya spp.) <strong>Forest</strong> Alliance (A.239)ELEMENT CONCEPTSummary: This basic dry-mesic forest of the southern Piedmont is dominated by Quercus alba and often Carya ovata incombination with other species of Quercus and Carya (i.e. Quercus stellata, Quercus prinus, Quercus velutina, Carya alba, Caryaglabra). It occurs on circumneutral to basic, well-drained soils in the Piedmont of Virginia, North Carolina, South Carolina, Georgia,and Alabama. Other canopy and subcanopy species that may occur include Liquidambar styraciflua, Carya ovalis, Liriodendrontulipifera, Juniperus virginiana var. virginiana, Cercis canadensis, Ulmus americana, Pinus taeda, and Pinus echinata. Proportionsof Liquidambar styraciflua and Acer rubrum increase following disturbance. Herbaceous species and vines that may occur within thiscommunity include Bignonia capreolata, Vitis rotundifolia, Chimaphila maculata, Asplenium platyneuron, Berchemia scandens,Carex nigromarginata, Carex albicans var. albicans, Polystichum acrostichoides, Botrychium virginianum, Tiarella cordifolia var.collina, and Goodyera repens. It is differentiated from non-basic oak-hickory forests by lacking such species as Quercus falcata andOxydendrum arboreum.Environment: This basic dry-mesic forest occurs on circumneutral to basic, well-drained soils in the southern Piedmont. A basic ormafic substrate is necessary for best development of this community and soils generally are in the order Alfisol. It commonly occurson mid to low slopes where conditions are somewhat ameliorated over upper slopes, i.e., in dry-mesic as opposed to dry conditions,but can occur in most topographic positions with circumneutral to basic soils that are well-drained.Vegetation: Stands of this forest are dominated by Quercus alba and often Carya ovata, in combination with other species ofQuercus and Carya (i.e., Quercus stellata, Quercus prinus, Quercus velutina, Carya alba, Carya glabra). Other canopy andsubcanopy species that may occur include Liquidambar styraciflua, Carya ovalis, Liriodendron tulipifera, Juniperus virginiana var.virginiana, Cercis canadensis, Ulmus americana, Pinus taeda, and Pinus echinata. Proportions of Liquidambar styraciflua and Acerrubrum increase following disturbance. Herbaceous species and vines that may occur within this community include Bignoniacapreolata, Vitis rotundifolia, Chimaphila maculata, Asplenium platyneuron, Berchemia scandens, Carex nigromarginata, Carexalbicans var. albicans, Polystichum acrostichoides, Botrychium virginianum, Tiarella cordifolia var. collina, and Goodyera repens.Community occurrences will have closed canopies and moderately well-developed subcanopy, shrub and herbaceous layers usuallyVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>63
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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• Maritime Oak - Holly Forest / W
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- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
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- Page 119 and 120: USFS Ecoregions: 221Ba:CCC, 221Ha:C
- Page 121 and 122: Classification Comments: On the Ban
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
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- Page 131 and 132: Lea, C. 2002a. Vegetation classific
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Tobe, J. D., J. E. Fairey, III, and