USFS Ecoregions: 212Fa:CCP, 212Fb:CCC, 212Ga:CCP, 212Gb:CCP, 221Am:CCP, 221Ba:CCC, 221Bd:CCC, 221Da:CCC,221Db:CCC, 221Ec:CCC, 221Ed:CCP, 221Ef:CCP, 221Ha:CCC, 221Hb:CCC, 221Hc:CCC, 221He:CCC, 222Eb:CCC, 222Ej:CCP,222En:CCC, 222Eo:CCC, 222Ha:CCC, 231Af:CCC, 231B:CC, 231Cd:CCC, 231D:CC, M221Aa:CCC, M221Ac:CCC,M221Ad:CCC, M221Bb:CCC, M221Bd:CCC, M221Be:CCC, M221Bf:CCC, M221Cd:CCC, M221Da:CCC, M221Dc:CCC,M222A:CC, M231A:CCFederal Lands: NPS (Little River Canyon?, Natchez Trace, Stones River); USFS (Bankhead, Cherokee, Daniel Boone, Oconee?,Ouachita, Ozark, Pisgah, <strong>Sumter</strong>?, Uwharrie)ELEMENT SOURCESReferences: Allard 1990, Anderson 1982, Anderson 1996, Fike 1999, Fleming et al. 2001, Hoagland 1997, Hoagland 2000, Major etal. 1999, McCoy 1958, Nelson 1986, Palmer-Ball et al. 1988, Peet et al. unpubl. data 2002, Penfound 1953, Schafale 1998b, Schafaleand Weakley 1990, Schmalzer and DeSelm 1982, Southeastern Ecology Working Group n.d.LOBLOLLY PINE - TULIPTREE / NORTHERN SPICEBUSH / FRINGED SEDGE FORESTELEMENT IDENTIFIERSNVC association: Pinus taeda - Liriodendron tulipifera / Lindera benzoin / Carex crinita <strong>Forest</strong>Database Code: CEGL007546Formation: Temporarily flooded mixed needle-leaved evergreen - cold-deciduous forest (I.C.3.N.b)Alliance: Pinus taeda - Liriodendron tulipifera Temporarily Flooded <strong>Forest</strong> Alliance (A.434)ELEMENT CONCEPTSummary: This broadly defined, successional wetland forest is dominated by Pinus taeda and Liriodendron tulipifera, but manyother canopy species are usually present. Lindera benzoin is a typical shrub, and Carex crinita is a typical herb. More information isneeded on the detailed floristics of this association. It develops in river floodplain alluvial terraces along streams following majordisturbances such as blowdowns, logging, and agriculture.Environment: This broadly defined, successional wetland forest develops in river floodplain alluvial terraces along streamsfollowing major disturbances such as blowdowns, logging, and agriculture. This is primarily a Zone IV community with a likelihoodof 51-100% of flooding with intermittent periodicity for 1-2 months (12.5-25% of total) of the growing season. Flooding usuallyoccurs in the spring. Just as important as flooding regime in determining the species composition of this community is land-usehistory since this community most often develops on floodplains subjected to timbering practices.Vegetation: This successional vegetation is dominated by Pinus taeda and Liriodendron tulipifera, but many other canopy speciesare usually present. Lindera benzoin is a typical shrub, and Carex crinita is a typical herb. More information is needed on the detailedfloristics of this association.Other woody species that may be present include Liquidambar styraciflua, Platanus occidentalis, Cornus florida, Acer rubrum, Nyssabiflora, Quercus alba, Fagus grandifolia, Morus rubra, Betula nigra, Fraxinus americana, Ilex opaca, Lindera benzoin, and Asiminaparviflora. Woody species common in the Piedmont portion of the range include Aesculus sylvatica and Acer negundo. Herbaceousand woody vine species that may occur include Carex crinita, Botrychium virginianum, Eupatorium purpureum, Polystichumacrostichoides, Arundinaria gigantea, Toxicodendron radicans ssp. radicans, Asplenium platyneuron, Woodwardia areolata,Osmunda cinnamomea, Ophioglossum vulgatum, Actaea racemosa (= Cimicifuga racemosa), Dulichium arundinaceum, Carexalbolutescens, Hypericum nudiflorum, Corydalis flavula, and others. The exotics Ligustrum sinense and Lonicera japonica commonlyoccur in forests of this alliance. The preceding information is from the related alliance description (Weakley et al. 1998), and thesefloristics were included in a 1995 CCA for the 'Pinus taeda - Liriodendron tulipifera / Lindera benzoin / Carex crinita Successional<strong>Forest</strong>' (1C03B020.01F), on which this association is apparently based. It is not clear where these data originated. Some of the taxamentioned (e.g., Eupatorium purpureum) seem "suspicious" and out of place.Apparently related vegetation from the Savannah River Site cited in Jones et al. (1981b) indicates dominance of some stands by Pinustaeda, Nyssa biflora, and Liriodendron tulipifera in the canopy, with Persea palustris, Magnolia virginiana, Acer rubrum, and Nyssabiflora in the subcanopy. Other species present include Vaccinium formosum, Viburnum nudum, Arundinaria gigantea,Toxicodendron radicans ssp. radicans, Mitchella repens, Osmunda cinnamomea, and Onoclea sensibilis.Dynamics: No informationSimilar Associations:• Liquidambar styraciflua - Liriodendron tulipifera / Onoclea sensibilis <strong>Forest</strong> (CEGL007329) -- a related deciduous type.Related Concepts:• Bottomland Hardwood - Yellow Pine (46) (USFS 1988) ?• IA8c. Lowland Pine - Oak <strong>Forest</strong> (Allard 1990) ?• Loblolly Pine - Hardwood: 82 (Eyre 1980) B• Loblolly pine - Swamp gum - Naked withe-rod Community (Jones et al. 1981b) ?• Loblolly pine-bottomland hardwood (Jones et al. 1981a) ?Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>98
Classification Comments: On the Bankhead <strong>National</strong> <strong>Forest</strong>, this community was observed on previously farmed alluvial terracesalong medium-sized streams. The canopy of this forest is dominated by Pinus taeda with Liriodendron tulipifera, Liquidambarstyraciflua, and Nyssa sylvatica also present in the canopy. Midstory components include Nyssa sylvatica, Magnolia macrophylla,Carpinus caroliniana, and Ostrya virginiana. Shrubs include Hamamelis virginiana, Lindera benzoin, and Asimina parviflora. Joneset al. (1981a) describe an old-growth stand at the Boiling Springs Natural Area at the DOE Savannah River Plant (upper coastal plain,Barnwell County, South Carolina) as a "loblolly pine-bottomland hardwood stand" which has a "senescent upperstory" composed ofPinus taeda and Liriodendron tulipifera. In contrast, the analysis of common forest types at the Savannah River Plant by Jones et al.(1981b) and Jones and Churchill (1987) includes floodplain vegetation dominated by Pinus taeda and Liquidambar styraciflua.CONSERVATION RANKING & RARE SPECIESGRank: GNA (modified/managed) (2002-8-19): This is a successional forest composed of species native to the southeastern UnitedStates; it is not of conservation concern and does not receive a conservation status rank.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This association is known from the Cumberland Plateau, Piedmont, South Atlantic Coastal Plain, and Chesapeake BayLowlands. It is also probably found in the East Gulf Coastal Plain and Upper East Gulf Coastal Plain.Subnations: AL, GA, NC, SC, VATNC Ecoregions: 43:P, 50:C, 52:C, 53:P, 56:C, 57:P, 58:?USFS Ecoregions: 231Aa:CCC, 231Ae:CCC, 231Af:CCC, 231An:CCC, 231Ao:CCP, 231Cd:CCC, 232Br:CCCFederal Lands: DOE (Savannah River Site); NPS (Kennesaw Mountain, Little River Canyon?); USFS (Bankhead, Oconee?,<strong>Sumter</strong>, Uwharrie)ELEMENT SOURCESReferences: Allard 1990, Eyre 1980, Jones and Churchill 1987, Jones et al. 1981a, Jones et al. 1981b, <strong>NatureServe</strong> Ecology -Southeastern U.S. unpubl. data, Nelson 1986, Peet et al. unpubl. data 2002, Southeastern Ecology Working Group n.d., USFS 1988,Weakley et al. 1998RIVERWEED HERBACEOUS VEGETATIONELEMENT IDENTIFIERSNVC association: Podostemum ceratophyllum Herbaceous VegetationDatabase Code: CEGL004331Formation: Permanently flooded temperate or subpolar hydromorphic-rooted vegetation (V.C.2.N.a)Alliance: Podostemum ceratophyllum Permanently Flooded Herbaceous Alliance (A.1752)ELEMENT CONCEPTSummary: This association represents essentially monospecific beds of Podostemum ceratophyllum, although some algae may alsobe present. This submersed vegetation forms a low mat or crust attached to rocks in moderately fast- to fast-flowing water. Typicalhabitat for this association in the Central Appalachians and related areas includes rocky surfaces of streambeds and riverbeds inmature drainage systems where the streams have cut down to rock and the floodplain is relatively narrow, or on dams. It tends to beassociated with higher pH streams which cut through diabase, limestone or calcareous shales.Environment: This association includes vegetation of shoals in rocky streambeds and riverbeds in mature drainage systems wherethe streams have cut down to rock, and the floodplain is relatively narrow; or on dams in moderately fast- to fast-flowing water. Ittends to be associated with higher pH streams which cut through diabase, limestone or calcareous shales. In the Piedmont region andpossibly elsewhere, the occurrence of macrophytic vegetation may be limited by unstable sediments, moderate to high gradients, andlarge variations in stream flow (Mulholland and Lenat 1992). Podostemum is one of the few vascular plants present in Piedmontstreams and rivers.Vegetation: This vegetation is almost always a monospecific community dominated by Podostemum ceratophyllum with no othervascular plants present; some Rhodophyta (red algae) may be present.Dynamics: Podostemum appears to require relatively stable streamflow to accumulate significant biomass; at high nutrient levelsfilamentous green algae may outcompete it (Mulholland and Lenat 1992). Dense beds of this vegetation may serve as importantsubstratum for a variety of invertebrates and other aquatic species (Nelson and Scott 1962). At least one rare species of snail of thePiedmont drainages, Somatogyrus virginicus, appears to be very closely tied to the occurrence of good beds of Podostemum (B.Adams pers. comm.). Although it is not clear whether the snail is actually dependent on Podostemum, it could be that Podostemum isjust a good habitat indicator.Similar Associations:Related Concepts:• Mountain river (Wharton 1978) ?• Open Water/Aquatic Bed Veg., Mountain Stream (Ambrose 1990a) B• River-weed shallow shore (CAP pers. comm. 1998) ?Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>99
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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• Maritime Oak - Holly Forest / W
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ALLIANCE CONCEPTSummary: This allia
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Related Concepts:• Yellow-Poplar:
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virginiana, Sanicula canadensis, De
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Federal Lands: COE (Claiborne Lake)
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ALLIANCE SOURCESReferences: Allard
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Quercus alba and Quercus velutina a
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USFS Ecoregions: 221Hc:CCC, 221He:C
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ALLIANCE CONCEPTSummary: This allia
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USFS Ecoregions: 221Eb:PP?, M221Ab:
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Federal Lands: NPS (Great Smoky Mou
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the associated species. More Appala
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ALLIANCE DISTRIBUTIONRange: This al
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USFS Ecoregions: 221Ha:CCC, 221Hc:C
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elevation environments such as Leio
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CULTIVATED FORESTEASTERN WHITE PINE
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Federal Lands: DOD (Arnold, Fort Be
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does not occur on wet soils. It occ
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Liriodendron tulipifera, although o
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
- Page 129 and 130: Gettman, R. W. 1974. A floristic su
- Page 131 and 132: Lea, C. 2002a. Vegetation classific
- Page 133 and 134: Pittman, Dr. Albert. Personal commu
- Page 135 and 136: Tobe, J. D., J. E. Fairey, III, and