fraseri, Menziesia pilosa, Heuchera villosa, Rhododendron catawbiense, Saxifraga michauxii, Sorbus americana, Oclemenaacuminata (= Aster acuminatus), Solidago glomerata. Other characteristic species are Minuartia groenlandica and Polypodiumappalachianum. On rock outcrops of felsic Anakeesta slate in the Great Smoky Mountains (from 1646-1987 m), typical speciesinclude Saxifraga michauxii, Carex misera, Calamagrostis cainii, Rhododendron carolinianum, Solidago glomerata, Oclemenaacuminata, Abies fraseri, and Leiophyllum buxifolium. Other characteristic species are Gentiana linearis and Calamagrostis cainii. Atlow to middle elevations (1256-1713 m) in the southern Appalachians on outcrops of mafic rock, or on felsic rock where perennialseepage exists, the sparse vegetation consists of graminoids, forbs, and shrubs. It is surrounded by deciduous forests dominated byQuercus rubra, Acer rubrum var. rubrum, and occasionally Tsuga caroliniana. Typical species here include Saxifraga michauxii,Coreopsis major, Schizachyrium scoparium, Kalmia latifolia, Dichanthelium acuminatum, Danthonia spicata, and Paronychiaargyrocoma. Other characteristic species include Campanula divaricata, Solidago bicolor, and Allium cernuum (= Alliumallegheniense). On amphibolite, metabasalt, metagabbro, or metagraywacke bedrock from 1350-1870 m elevation within a matrix ofQuercus rubra <strong>Forest</strong> or high elevation grasslands and shrublands, the vegetation includes Saxifraga michauxii, Danthonia spicata,Krigia montana, Carex misera, Angelica triquinata, Athyrium filix-femina ssp. asplenioides, Rhododendron catawbiense, andHeuchera villosa. Other characteristic species are Sanguisorba canadensis, Sibbaldiopsis tridentata, Hylotelephium telephioides (=Sedum telephioides), Houstonia purpurea var. montana, Geum radiatum, Solidago spithamaea, and Huperzia appalachiana. In thecentral Blue Ridge mountains of Virginia, at elevations of 850-1200 m, this alliance occurs on greenstone (metabasalt, a maficmetamorphic rock). Characteristic herbaceous species include Hylotelephium telephioides, Solidago simplex var. randii, Heucherapubescens, Deschampsia flexuosa, Houstonia longifolia (= Houstonia longifolia var. compacta), Dennstaedtia punctilobula,Campanula divaricata, Agrostis perennans, Carex pensylvanica, Saxifraga michauxii, Arabis lyrata, Allium allegheniense?, Phloxsubulata ssp. brittonii, Sibbaldiopsis tridentata, Liatris turgida, Huperzia appalachiana, Polypodium appalachianum, Gymnocarpiumappalachianum, and Oclemena acuminata. Shrubs include Physocarpus opulifolius, Sorbus americana, Betula alleghaniensis,Quercus rubra, Ribes rotundifolium, Diervilla lonicera, Hamamelis virginiana, Ilex montana, Kalmia latifolia, Menziesia pilosa, andAbies balsamea. Rare alpine disjunct species are sometimes present, including Juncus trifidus and Trisetum spicatum.Related Concepts:• Aster acuminatus / Menziesia pilosa outcrop community (Wiser et al. 1996) ?• Aster acuminatus / Menziesia pilosa outcrop community (Wiser 1993) ?• Calamagrostis cainii / Rhododendron carolinianum outcrop community (Wiser et al. 1996) ?• Calamagrostis cainii / Rhododendron carolinianum outcrop community (Wiser 1993) ?• Coreopsis major / Schizachyrium scoparium outcrop community (Wiser et al. 1996) ?• Coreopsis major / Schizachyrium scoparium outcrop community (Wiser 1993) ?• Deschampsia flexuosa / Angelica triquinata outcrop community (Wiser et al. 1996) ?• Deschampsia flexuosa / Angelica triquinata outcrop community (Wiser 1993) ?• Paronychia argycoma (sic) / Polypodium appalachianum outcrop community (Wiser et al. 1996) ?• Paronychia argycoma (sic) / Polypodium appalachianum outcrop community (Wiser 1993) ?• Paronychia argyrocoma - Potentilla tridentata - Arenaria groenlandica Association (Rawinski and Wieboldt 1993) ?• High Elevation Rocky Summit (Schafale and Weakley 1990) I• IE4a. Southern Appalachian High Elevation Acidic Rocky Summit (Allard 1990) IClassification Comments: NoneALLIANCE DISTRIBUTIONRange: This alliance is found in the southern and central Blue Ridge (metamorphic rock portions of the southern and centralAppalachians), in Georgia, North Carolina, South Carolina, Tennessee, and Virginia.Subnations: GA, NC, SC, TN, VATNC Ecoregions: 51:C, 52:C, 59:CUSFS Ecoregions: M221Aa:CCP, M221Ab:CCC, M221Ad:CCC, M221Da:CCC, M221Db:CCC, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Great Smoky Mountains, Shenandoah); USFS (Chattahoochee, Cherokee, George Washington, Nantahala,Pisgah, <strong>Sumter</strong>)ALLIANCE SOURCESReferences: Allard 1990, Feldcamp 1984, Rawinski and Wieboldt 1993, Schafale and Weakley 1990, Wiser 1993, Wiser et al. 1996V.B.2.N.d. Temporarily flooded temperate perennial forb vegetationV.B.2.N.D. JUSTICIA AMERICANA TEMPORARILY FLOODED HERBACEOUS ALLIANCE(A.1657)COMMON WATER-WILLOW TEMPORARILY FLOODED HERBACEOUS ALLIANCEALLIANCE CONCEPTSummary: This alliance covers rocky river shoals dominated by Justicia americana with Orontium aquaticum, Podostemumceratophyllum, Leersia spp., Lemna minor, Saururus cernuus, and others. A sparse canopy may be present, and species may includeCarpinus caroliniana ssp. caroliniana, Fagus grandifolia, and Fraxinus pennsylvanica. There is some apparent regional variation inVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>30
the associated species. More Appalachian examples may contain Orontium aquaticum as a codominant. In parts of the Ridge andValley and Piedmont, Hymenocallis caroliniana (= Hymenocallis coronaria) is codominant. In the Edwards Plateau of central Texas,associated with Justicia americana are Bacopa monnieri, Fuirena simplex, Eleocharis geniculata (= Eleocharis caribaea), Eleocharismontevidensis, and Cyperus spp.Related Concepts:• Justicia americana herbaceous alliance (Hoagland 2000) ?• IIE3a. Riverside Shoal and Stream Bar Complex (Allard 1990) I Rocky Bar and Shore (Schafale and Weakley 1990) I• Shoal and Stream Bar (Nelson 1986) I• Water-willow (Justicia americana) - smart-weed riverbed community (Fike 1999) ?Classification Comments: NoneALLIANCE DISTRIBUTIONRange: This alliance is distributed in the Edwards Plateau of Texas, Ozark Highlands, Boston Mountains, Ouachita Mountains,Interior Low Plateau, Cumberland Plateau, Piedmont, and Arkansas Valley. It is found in Ohio, Alabama, Arkansas, Georgia, NorthCarolina, Oklahoma, South Carolina, Tennessee, Texas, and possibly Virginia (?).Subnations: AL, AR, GA, KY, MD?, NC, OH, OK, PA, SC, TN, TX, VA?, WVTNC Ecoregions: 29:C, 38:C, 39:C, 43:C, 44:C, 45:C, 48:C, 49:C, 50:C, 51:C, 52:C, 59:C, 60:C, 61:CUSFS Ecoregions: 212Fa:CCP, 212Fb:CCC, 212Ga:CCP, 212Gb:CCP, 221Am:CCP, 221Ba:CCC, 221Bd:CCC, 221Da:CCC,221Db:CCC, 221Ec:CCC, 221Ed:CCP, 221Ef:CCP, 221Ha:CCC, 221Hb:CCC, 221Hc:CCC, 221He:CCC, 222Ab:CCC, 222Ag:CCC,222Ah:CCC, 222An:CCC, 222Eb:CCC, 222Eg:CCC, 222Ej:CCP, 222En:CCC, 222Eo:CCC, 222Ha:CCC, 231Af:CCC, 231B:CC,231Cd:CCC, 231Dc:CCC, 231Ga:CCC, 231Gb:CCC, 231Gc:CCC, 315D:CC, 321B:PP, M221Aa:CCC, M221Ac:CCC,M221Ad:CCC, M221Bb:CCC, M221Bd:CCC, M221Be:CCC, M221Bf:CCC, M221Cd:CCC, M221Da:CCC, M221Dc:CCC,M222Aa:CCC, M222Ab:CCC, M231Aa:CCC, M231Ab:CCC, M231Ac:CCC, M231Ad:CCCFederal Lands: NPS (Little River Canyon?, Natchez Trace, Stones River); USFS (Bankhead, Cherokee, Daniel Boone, Oconee?,Ouachita, Ozark, Pisgah, <strong>Sumter</strong>, Uwharrie); USFWS (Cahaba River)ALLIANCE SOURCESReferences: Allard 1990, Faber-Langendoen et al. 1996, Fike 1999, Foti et al. 1994, Hoagland 2000, Nelson 1986, Schafale andWeakley 1990, Schmalzer and DeSelm 1982V.B.2.N.f. Saturated temperate perennial forb vegetationV.B.2.N.F. VITTARIA APPALACHIANA - HEUCHERA PARVIFLORA SATURATED HERBACEOUSALLIANCE (A.1696)APPALACHIAN SHOESTRING FERN - CAVE ALUMROOT SATURATED HERBACEOUSALLIANCEALLIANCE CONCEPTSummary: This alliance accommodates saturated communities associated with overhanging to vertical rocks and a seasonal toperennial waterfall as a source of aqueous aerosol. One association includes sparse to moderately dense vegetation of sandstonerockhouses in portions of the Cumberland Plateau, where seasonal waterfalls and strongly overhanging erosion features calledrockhouses provide moist conditions. Vittaria appalachiana and Heuchera parviflora are dominant and characteristic. Endemics suchas Ageratina luciae-brauniae and Solidago albopilosa are associated with the floors of the rockhouses and often dominate thevegetative cover, and the endemic Minuartia cumberlandensis also sometimes occupies the floor but is more characteristic ofsomewhat drier sandstone exposures. Thalictrum mirabile is also endemic to this community but is more characteristic of seepages onthe rockhouse walls. In the Southern Blue Ridge escarpment region, this alliance includes herbaceous vegetation on rock substratesassociated with waterfalls, on nearly vertical rock surfaces and ledges, slopes, and crevices with shallow soils which are constantlysaturated. Other characteristic species include Huperzia porophila, Asplenium montanum, Asplenium trichomanes ssp. trichomanes,Asplenium monanthes, Cystopteris protrusa, Polypodium virginianum, Trichomanes boschianum, Grammitis nimbata (=Micropolypodium nimbatum), Hymenophyllum tayloriae, Trichomanes intricatum, Phegopteris connectilis, Adiantum pedatum,Saxifraga careyana, Saxifraga caroliniana, Impatiens capensis, Hydrocotyle americana, Thalictrum spp., Oxalis montana, Carexbiltmoreana, Galax urceolata, Sphagnum quinquefarium, Sphagnum girgensohnii, Plagiomnium carolinianum, Plagiomnium affine,Mnium marginatum, Pseudotaxiphyllum distichaceum, Bryocrumia vivicolor, Thamnobryum alleghaniense, Oncophorus raui,Hyophila involuta, Dichodontium pellucidum, Plagiochila sharpii ssp. sharpii, Plagiochila caduciloba, Plagiochila sullivantii,Plagiochila austini, Fissidens osmundioides, Bazzania denudata, Conocephalum conicum, Pellia epiphylla, Pellia neesiana, andRiccardia multifida.Related Concepts:• Cumberland Plateau sandstone glade (Evans 1991) I• IID5a. Wet Acidic Cliff (Allard 1990) I• Moist sandstone cliff (Evans 1991) I• Spray Cliff (Nelson 1986) IVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 200431Copyright © 2004 <strong>NatureServe</strong>
- Page 3: 1 NatureServe is an international o
- Page 11 and 12: types. However, when necessary, mod
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- Page 15 and 16: Generally 5 or fewer occurrences an
- Page 17 and 18: All scientific names for vascular s
- Page 19 and 20: Summary: A short description of the
- Page 21 and 22: Loucks, O. 1996. 100 years after Co
- Page 23 and 24: ALLIANCES BY US NATIONAL VEGETATION
- Page 25 and 26: virginiana, and Acer pensylvanicum
- Page 27 and 28: virginiana, Juniperus virginiana va
- Page 29 and 30: • Appalachian pine-oak forest (Ev
- Page 31 and 32: • Maritime Oak - Holly Forest / W
- Page 33 and 34: ALLIANCE CONCEPTSummary: This allia
- Page 35 and 36: Related Concepts:• Yellow-Poplar:
- Page 37 and 38: virginiana, Sanicula canadensis, De
- Page 39 and 40: Federal Lands: COE (Claiborne Lake)
- Page 41 and 42: ALLIANCE SOURCESReferences: Allard
- Page 43 and 44: Quercus alba and Quercus velutina a
- Page 45 and 46: USFS Ecoregions: 221Hc:CCC, 221He:C
- Page 47 and 48: ALLIANCE CONCEPTSummary: This allia
- Page 49 and 50: USFS Ecoregions: 221Eb:PP?, M221Ab:
- Page 51: Federal Lands: NPS (Great Smoky Mou
- Page 55 and 56: ALLIANCE DISTRIBUTIONRange: This al
- Page 57 and 58: USFS Ecoregions: 221Ha:CCC, 221Hc:C
- Page 59 and 60: elevation environments such as Leio
- Page 61 and 62: CULTIVATED FORESTEASTERN WHITE PINE
- Page 63 and 64: Federal Lands: DOD (Arnold, Fort Be
- Page 65 and 66: does not occur on wet soils. It occ
- Page 67 and 68: Liriodendron tulipifera, although o
- Page 69 and 70: ELEMENT SOURCESReferences: Allard 1
- Page 71 and 72: ELEMENT SOURCESReferences: Allard 1
- Page 73 and 74: Similar Associations:• Pinus echi
- Page 75 and 76: • Pinus echinata - Quercus stella
- Page 77 and 78: this association occurs in edaphica
- Page 79 and 80: Vegetation: This forest typically h
- Page 81 and 82: strobus can have high coverage and
- Page 83 and 84: CONSERVATION RANKING & RARE SPECIES
- Page 85 and 86: • Quercus alba - Carya alba / Euo
- Page 87 and 88: muehlenbergii. In addition, Acer ba
- Page 89 and 90: WHITE OAK - MOCKERNUT HICKORY / AME
- Page 91 and 92: CONSERVATION RANKING & RARE SPECIES
- Page 93 and 94: Vegetation: This association encomp
- Page 95 and 96: ELEMENT DISTRIBUTIONRange: This com
- Page 97 and 98: • Quercus prinus - Carya spp. - Q
- Page 99 and 100: catesbaei, Desmodium nudiflorum, Eu
- Page 101 and 102: ELEMENT SOURCESReferences: Allard 1
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High-ranked species: No information
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Dynamics: No informationSimilar Ass
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protrusa, Grammitis nimbata (= Micr
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ELEMENT DISTRIBUTIONRange: This for
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COMMON WATER-WILLOW HERBACEOUS VEGE
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• Betula nigra - Platanus occiden
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Formation: Temporarily flooded temp
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epeated flooding, this community ma
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USFS Ecoregions: 221Ba:CCC, 221Ha:C
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Classification Comments: On the Ban
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Classification Comments: This veget
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BIBLIOGRAPHYALNHP [Alabama Natural
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DeYoung, H. R. 1979. The white pine
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Gettman, R. W. 1974. A floristic su
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Lea, C. 2002a. Vegetation classific
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Pittman, Dr. Albert. Personal commu
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Tobe, J. D., J. E. Fairey, III, and