Alliance: Carex torta Temporarily Flooded Herbaceous Alliance (A.1340)ELEMENT CONCEPTSummary: This association accommodates Carex torta-dominated alluvial wetlands on sand, gravel, and rock bars along riverbanksin valleys and gorges in the southern Appalachians, ranging west into the Cumberland Plateau and the Interior Low Plateau, and northinto the Central Appalachians. This association is characterized by light-requiring, tough-rooted herbaceous perennials tolerant offrequent inundation and flood-scouring. Carex torta often forms dense, extensive colonies. Associated species vary with geographybut can include Verbena hastata, Doellingeria umbellata (= Aster umbellatus), Dichanthelium clandestinum, Solidago rugosa ssp.aspera, Juncus effusus, Scirpus expansus, Scirpus cyperinus, Equisetum arvense, Onoclea sensibilis, Vernonia noveboracensis,Lycopus virginicus, Scutellaria lateriflora, Lobelia cardinalis, Symphyotrichum dumosum (= Aster dumosus), Lycopus virginicus,Osmunda regalis, Hypericum mutilum, Eupatorium fistulosum, Solidago patula, and Salix sericea.Environment: This association occupies boulder and gravel bars on the frequently flooded, active channel shelves of high-gradientrivers and large streams. Along Laurel Fork, flooding duration is probably similar to that documented along Passage Creek inShenandoah County, Virginia, by Hupp (1982). In that drainage, the channel shelf was inundated approximately 15% of the time.Periodic large or severe floods transport and deposit large numbers of cobbles, stones, and even boulders in characteristic bars bothwithin the channel (islands) and along its edges (streambanks). These bars provide a matrix for deposition of finer alluvium andhabitats for the establishment of tough, adaptable herbaceous plants, which in turn stabilize the bars with massive networks ofperennial rootstocks. Regular flood-scouring batters or removes woody plants which take root in these habitats, maintaining opencanopyconditions. These habitats are highly dynamic, with conditions more or less constantly shifting in response to an irregular butpowerful disturbance regime. While some of the bar habitats may be damaged or removed by severe floods, others may accrete or benewly deposited during the same events.Vegetation: Vegetation is characterized by light-demanding, tough-rooted herbaceous perennials tolerant of frequent inundation andflood-scouring. Carex torta is the dominant species and typically forms dense, extensive colonies. Associated species vary withgeography but can include Verbena hastata, Doellingeria umbellata (= Aster umbellatus), Dichanthelium clandestinum, Solidagorugosa ssp. aspera, Juncus effusus var. solutus, Scirpus expansus, Scirpus cyperinus, Equisetum arvense, Onoclea sensibilis,Vernonia noveboracensis, Lycopus virginicus, Scutellaria lateriflora, Lobelia cardinalis, Symphyotrichum dumosum (= Asterdumosus), Osmunda regalis var. spectabilis, Eupatorium fistulosum, Solidago patula, and Hypericum mutilum. Small specimens ofSalix sericea and other woody plants are also frequent.Dynamics: Periodic large or severe floods transport and deposit large numbers of cobbles, stones, and even boulders in characteristicbars both within the channel (islands) and along its edges (streambanks). These bars provide a matrix for deposition of finer alluviumand habitats for the establishment of tough, adaptable herbaceous plants, which in turn stabilize the bars with massive networks ofperennial rootstocks. Regular flood-scouring batters or removes woody plants which take root in these habitats, maintaining opencanopyconditions. These habitats are highly dynamic, with conditions more or less constantly shifting in response to an irregular butpowerful disturbance regime. While some of the bar habitats may be damaged or removed by severe floods, others may accrete or benewly deposited during the same events (Hupp 1982). Successionally, this unit can be considered a pioneering type on new, coarsealluvial land, but it is also more or less permanently maintained by natural disturbances.Similar Associations:• Carex torta - Apocynum cannabinum Herbaceous Vegetation (CEGL006536)Related Concepts:• Carex torta Association (Fleming and Moorhead 1996) ?• Carex torta Herbaceous Vegetation (Fleming and Coulling 2001) ?• IIE3a. Riverside Shoal and Stream Bar Complex (Allard 1990) B• Riverine Upper and Lower Perennial Beach/Bar, Cobble/Gravel, Seasonally Flooded (R2and3BB1C) (Cowardin et al. 1979) ?• Riverine Upper and Lower Perennial Beach/Bar, Sand, Seasonally Flooded (R2and3BB2C) (Cowardin et al. 1979) ?• Rocky Bar and Shore (Twisted Sedge Subtype) (Schafale 1998b) ?• Torturous sedge gravel rivershore (CAP pers. comm. 1998) ?Classification Comments:CONSERVATION RANKING & RARE SPECIESGRank: G3G4 (2000-1-21): This community is found as linear occurrences along waterways in the Southern Appalachians, westinto the Cumberland Plateau and the Interior Low Plateau, and north into the Central Appalachians and Allegheny Mountains.Although this community has a moderately large geographic range, it is uncommon within its range and occurrences are small. Thiscommunity is more common than the number of documented occurrences would suggest, since it is often overlooked in inventories.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This community is found as linear occurrences along waterways in the Southern Appalachians, west into the CumberlandPlateau and the Interior Low Plateau, and north into the Central Appalachians, and Allegheny Mountains.Subnations: AL, DE, GA, KY, NC, SC, TN, VA, WVTNC Ecoregions: 44:C, 50:C, 51:C, 58:C, 59:CVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>96
USFS Ecoregions: 221Ba:CCC, 221Ha:CC?, 221Hb:CCC, 221Hc:CCC, 221He:CCC, 222Eg:CCC, 222En:CCC, 222Eo:CCC,231Cd:CCC, M221Aa:CCC, M221Ab:CCC, M221Ac:CCC, M221Ad:CCP, M221Ba:CCC, M221Bb:CCC, M221Bd:CCC,M221Be:CCC, M221Cd:CCP, M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Great Smoky Mountains); USFS (Bankhead, Chattahoochee, Cherokee, Daniel Boone, George Washington,Jefferson, Nantahala, Pisgah, <strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, CAP pers. comm. 1998, Cowardin et al. 1979, Eyre 1980, Fleming and Coulling 2001, Fleming andMoorhead 1996, Fleming et al. 2001, Hupp 1982, <strong>NatureServe</strong> Ecology - Southeastern U.S. unpubl. data, Palmer-Ball et al. 1988,Peet et al. unpubl. data 2002, Schafale 1998b, Schafale and Weakley 1990, Schafale pers. comm., Southeastern Ecology WorkingGroup n.d.COMMON WATER-WILLOW HERBACEOUS VEGETATIONSOUTHERN PIEDMONT SMALL FLOODPLAIN AND RIPARIAN FORESTELEMENT IDENTIFIERSNVC association: Justicia americana Herbaceous VegetationDatabase Code: CEGL004286Formation: Temporarily flooded temperate perennial forb vegetation (V.B.2.N.d)Alliance: Justicia americana Temporarily Flooded Herbaceous Alliance (A.1657)ELEMENT CONCEPTSummary: This association is found primarily in the Piedmont, Cumberland Plateau, Interior Low Plateau, Ozarks, OuachitaMountains, and adjacent provinces. Stands occur on the shoals or bars of rocky streams and riverbeds. It provides habitat in someportions of its range for globally rare dragonflies and herbs. Justicia americana is the characteristic dominant. Other herbaceousspecies that may be present include Diodia teres, Gratiola brevifolia, Leersia sp., Lemna minor, Orontium aquaticum, Podostemumceratophyllum, Scirpus sp., Saururus cernuus, and Xyris difformis var. difformis. A sparse canopy layer, which can include Carpinuscaroliniana ssp. caroliniana, Salix interior, Fagus grandifolia, and Fraxinus pennsylvanica among other species, may be present.Environment: This association occurs on the shoals or bars of rocky streams and riverbeds, or gravelly sands.Vegetation: Justicia americana is the characteristic dominant. Other herbaceous species that may be present include Diodia teres,Gratiola brevifolia, Leersia sp., Lemna minor, Orontium aquaticum, Podostemum ceratophyllum, Saururus cernuus, and Xyrisdifformis var. difformis. In Ohio, Justicia usually grows in nearly pure patches, so that few other species are associated with it. Bidensspp., Cuscuta gronovii, Mimulus ringens, Polygonum spp., Rumex spp., and Salix interior can occur (Anderson 1982). A sparsecanopy layer may be present, which can include Carpinus caroliniana, Fagus grandifolia, and Fraxinus pennsylvanica, among others.In the Cumberland Plateau of Alabama, Justicia americana is present in dense patches with some interspersion of other speciesincluding Pilea pumila, Boehmeria cylindrica, Eclipta prostrata (= Eclipta alba), Juncus coriaceus, Mikania scandens, Ludwigiapalustris, Leersia sp. and Bidens sp. Schmalzer and DeSelm (1982) discuss Orontium aquaticum growing along streambanks or inshallow riffles "along or with" Justicia americana in the Obed River in the Cumberland Plateau of Tennessee.Dynamics: Stands in some situations may be obliterated by ongoing river channeling. Anderson (1982) describes some of the lifehistorycharacteristics of Justicia americana that allow it to persist in river channels.Similar Associations:Related Concepts:• Aquatic Types (Schmalzer and DeSelm 1982) B• IIE3a. Riverside Shoal and Stream Bar Complex (Allard 1990) B• Rocky Bar and Shore (Water Willow Subtype) (Schafale 1998b) ?Classification Comments: This type, in Ohio, often forms pure patches, but consistent identification may require a simple cutoffrule, such as at least 50% cover of Justicia (Anderson 1982). However, Anderson (1996) no longer recognizes this type.CONSERVATION RANKING & RARE SPECIESGRank: G4G5 (1997-9-12): No informationHigh-ranked species: No informationELEMENT DISTRIBUTIONRange: This type is found primarily in the Piedmont, Interior Low Plateau, Cumberland Plateau, Ozarks, Ouachita Mountains, andadjacent provinces. It ranges from Alabama, Georgia and the Carolinas west to Arkansas and Oklahoma and north to Ohio,Pennsylvania, and Delaware.Subnations: AL, AR, GA, KY, MD?, NC, OH, OK, PA, SC?, TN, VA?, WVTNC Ecoregions: 38:C, 39:C, 43:C, 44:C, 45:C, 48:C, 49:C, 50:C, 51:C, 52:C, 59:C, 60:C, 61:CVegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>97
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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ALLIANCES BY US NATIONAL VEGETATION
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virginiana, and Acer pensylvanicum
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virginiana, Juniperus virginiana va
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• Appalachian pine-oak forest (Ev
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• Maritime Oak - Holly Forest / W
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ALLIANCE CONCEPTSummary: This allia
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Related Concepts:• Yellow-Poplar:
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virginiana, Sanicula canadensis, De
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Federal Lands: COE (Claiborne Lake)
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ALLIANCE SOURCESReferences: Allard
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Quercus alba and Quercus velutina a
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USFS Ecoregions: 221Hc:CCC, 221He:C
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ALLIANCE CONCEPTSummary: This allia
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USFS Ecoregions: 221Eb:PP?, M221Ab:
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Federal Lands: NPS (Great Smoky Mou
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the associated species. More Appala
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ALLIANCE DISTRIBUTIONRange: This al
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USFS Ecoregions: 221Ha:CCC, 221Hc:C
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elevation environments such as Leio
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CULTIVATED FORESTEASTERN WHITE PINE
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Federal Lands: DOD (Arnold, Fort Be
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does not occur on wet soils. It occ
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- Page 125 and 126: BIBLIOGRAPHYALNHP [Alabama Natural
- Page 127 and 128: DeYoung, H. R. 1979. The white pine
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