• Pinus echinata - Quercus (prinus, falcata) / Oxydendrum arboreum / Vaccinium pallidum <strong>Forest</strong> (CEGL007493)Related Concepts:• IA7a. Xeric Shortleaf Pine - Oak <strong>Forest</strong> (Allard 1990) BClassification Comments: Defined from the Chattooga Basin Project (S. Simon pers. comm.). It is unclear if this is a uniqueassociation, or if it would be better considered as a subtype of Pinus echinata - Quercus (prinus, falcata) / Oxydendrum arboreum /Vaccinium pallidum <strong>Forest</strong> (CEGL007493).CONSERVATION RANKING & RARE SPECIESGRank: G2G3 (1997-8-15): No informationHigh-ranked species: No informationELEMENT DISTRIBUTIONRange: This community occurs in the Blue Ridge/Piedmont transition of South Carolina and Georgia.Subnations: GA, SCTNC Ecoregions: 51:CUSFS Ecoregions: M221Dc:CCC, M221Dd:CCCFederal Lands: USFS (Chattahoochee, <strong>Sumter</strong>)ELEMENT SOURCESReferences: Allard 1990, <strong>NatureServe</strong> Ecology - Southeastern U.S. unpubl. data, Schafale and Weakley 1990, Simon pers. comm.,Southeastern Ecology Working Group n.d.SHORTLEAF PINE - WHITE OAK / HILLSIDE BLUEBERRY / ARROWLEAF HEARTLEAF - STRIPED WINTERGREENFORESTELEMENT IDENTIFIERSNVC association: Pinus echinata - Quercus alba / Vaccinium pallidum / Hexastylis arifolia - Chimaphila maculata <strong>Forest</strong>Database Code: CEGL008427Formation: Mixed needle-leaved evergreen - cold-deciduous forest (I.C.3.N.a)Alliance: Pinus echinata - Quercus (alba, falcata, stellata, velutina) <strong>Forest</strong> Alliance (A.394)ELEMENT CONCEPTSummary: This association includes forests dominated by a mixture of Pinus echinata and mesophytic and dry-mesophytic oaks(e.g., Quercus alba, Quercus rubra, Quercus velutina) occurring in the Piedmont of the southeastern United States, ranging north andwest through the Southern Ridge and Valley, Cumberland Plateau, low Southern Blue Ridge, and upper Piedmont. These forestsoccur on low to middle slope positions, on protected to intermediately exposed sites. The mixed evergreen - deciduous canopy isdominated by Pinus echinata and Quercus alba, sometimes with high coverage by other Quercus spp. (Quercus velutina, Quercuscoccinea, Quercus falcata, Quercus rubra). Xerophytic Quercus spp. such as Quercus prinus, Quercus stellata, as well as otherspecies of pines may be present, but are typically not abundant. A well-developed subcanopy is typical, with species such as Acerrubrum, Nyssa sylvatica, Carya glabra, Cornus florida, and Oxydendrum arboreum. The shrub stratum is sparse to patchy with lowshrubs (Vaccinium pallidum, Vaccinium stamineum, Vaccinium arboreum, Chimaphila maculata), and vines (Vitis rotundifolia). Theherb stratum is patchy to absent. Hexastylis arifolia is a typical herb. Stands without fire management may experience invasion byAcer rubrum. Piptochaetium avenaceum may be an important grass in more open stands.Environment: These forests occur on low to middle slope positions, on protected to intermediately exposed sites.Vegetation: The mixed evergreen - deciduous canopy of stands is dominated by Pinus echinata and Quercus alba, sometimes withhigh coverage by other Quercus spp. (Quercus velutina, Quercus coccinea, Quercus falcata, Quercus rubra). Xerophytic Quercusspp. such as Quercus prinus, Quercus stellata, as well as other species of pines may be present, but are typically not abundant. Awell-developed subcanopy is typical, with species such as Acer rubrum, Nyssa sylvatica, Carya glabra, Cornus florida, andOxydendrum arboreum. The shrub stratum is sparse to patchy with low shrubs (Vaccinium pallidum, Vaccinium stamineum,Vaccinium arboreum, Chimaphila maculata), and vines (Vitis rotundifolia). The herb stratum is patchy to absent. Hexastylis arifoliais a typical herb. Stands without fire management may experience invasion by Acer rubrum. Piptochaetium avenaceum may be animportant grass in more open stands. A dense forest from the Talladega <strong>National</strong> <strong>Forest</strong>, Talladega Ranger District, included here, isdominated by Quercus coccinea, Pinus echinata; other canopy components include Quercus velutina, Quercus alba, Quercus falcata,Liriodendron tulipifera, Pinus taeda, Carya glabra, and Liquidambar styraciflua. The patchy shrub layer includes Vacciniumarboreum, Vaccinium pallidum, Viburnum acerifolium, and Acer rubrum. The sparse herbaceous layer is characterized byPiptochaetium avenaceum, which may be an important grass in more open stands.Dynamics: No informationSimilar Associations:• Pinus echinata - Quercus (prinus, falcata) / Oxydendrum arboreum / Vaccinium pallidum <strong>Forest</strong> (CEGL007493)Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>52
• Pinus echinata - Quercus stellata - Quercus prinus - Carya glabra / (Danthonia spicata, Piptochaetium avenaceum) <strong>Forest</strong>(CEGL007500) -- is a more open, grassy variant.• Quercus alba - Quercus falcata / Vaccinium (arboreum, hirsutum, pallidum) <strong>Forest</strong> (CEGL008567) -- is a related, primarilydeciduous type of the Ridge and Valley and parts of the Southern Blue Ridge adjacent to the Ridge and Valley.• Quercus falcata - Quercus alba - Carya alba / Oxydendrum arboreum / Vaccinium stamineum <strong>Forest</strong> (CEGL007244) -- is arelated, primarily deciduous type with representation in the Piedmont and Ridge and Valley but not in the Blue Ridge.Related Concepts: No informationClassification Comments: This forest has an overall more mesophytic species composition and occurs on deeper soil or on moreprotected sites than the more extreme shortleaf pine - oak forest, Pinus echinata - Quercus (prinus, falcata) / Oxydendrum arboreum /Vaccinium pallidum <strong>Forest</strong> (CEGL007493). In the Daniel Boone <strong>National</strong> <strong>Forest</strong> (Kentucky) this vegetation is important as part of apine-oak matrix which is significant for restoration of Red-cockaded Woodpecker (Picoides borealis) habitat. Piptochaetiumavenaceum may be an important grass in more open stands. Some plots attributed to this type have more Quercus alba than Pinusechinata.CONSERVATION RANKING & RARE SPECIESGRank: G3G4 (2002-10-23): Although this association has a reasonably wide potential natural range, shortleaf pine (Pinusechinata) populations seem to have undergone rangewide declines in vigor and extent. This phenomenon is especially pronounced inthe range of this type, primarily due to changes in fire regime and to depredations of the Southern Pine Beetle (Dendroctonusfrontalis). This community has had little inventory, but the total acreage in viable condition is believed to be quite limited. The moremesic to submesic habitat of this association may never have been common and is likely more vulnerable to successional changes thanmore xeric stands. Further, stands of this association are threatened by removal of commercially valuable tree species (e.g., Quercusalba, Quercus rubra, Pinus echinata), as well as by conversion to commercial forest plantations, and by the effects of continued firesuppression, which would inhibit the reproduction of Pinus echinata and cause the grass-dominated herbaceous layer to deteriorate.Following the removal of the commercially valuable species, and in the absence of fire, stands could become populated withsuccessional hardwoods (e.g., Liriodendron tulipifera, Liquidambar styraciflua) as well as less fire-adapted pines (Pinus taeda, Pinusvirginiana). The range in the rank reflects the need for further inventory and evaluation of this community.High-ranked species: No informationELEMENT DISTRIBUTIONRange: This community occurs in the Piedmont of the southeastern United States, ranging north and west through the SouthernRidge and Valley, Cumberland Plateau, and low Southern Blue Ridge.Subnations: AL, GA, KY, NC, SC, TN, VA?TNC Ecoregions: 50:C, 51:C, 52:CUSFS Ecoregions: 221H:PP, 221I:PP, 221J:PP, 222E:PP, 231Ab:CCC, 231C:CP, 231Db:CCC, 231Dc:CCC, M221C:CP,M221Dc:CCC, M221Dd:CCCFederal Lands: NPS (Carl Sandburg Home, Kings Mountain, Little River Canyon?); USFS (Chattahoochee, Cherokee, DanielBoone, <strong>Sumter</strong>, Talladega)ELEMENT SOURCESReferences: <strong>NatureServe</strong> Ecology - Southeastern U.S. unpubl. data, Southeastern Ecology Working Group n.d.SHORTLEAF PINE / (HILLSIDE BLUEBERRY, DEERBERRY) - MOUNTAIN LAUREL FORESTELEMENT IDENTIFIERSNVC association: Pinus echinata / Vaccinium (pallidum, stamineum) - Kalmia latifolia <strong>Forest</strong>Database Code: CEGL007078Formation: Rounded-crowned temperate or subpolar needle-leaved evergreen forest (I.A.8.N.b)Alliance: Pinus echinata <strong>Forest</strong> Alliance (A.119)ELEMENT CONCEPTSummary: This association includes forest vegetation with greater than 75% of the canopy cover of Pinus echinata, occurring over ashrub stratum dominated by ericaceous species, typically Vaccinium pallidum, Vaccinium stamineum, and Kalmia latifolia. Deciduousspecies make-up less than 25% of the canopy coverage and may include Quercus falcata, Quercus coccinea, or, in the southern partof this association's range, Quercus stellata and Quercus marilandica. This community often has a midstory tree stratum withOxydendrum arboreum, Carya pallida, Cornus florida, or Diospyros virginiana. Other characteristic species include Smilax glauca,Silphium compositum, Pteridium aquilinum var. latiusculum, Scleria oligantha, Piptochaetium avenaceum, and Tephrosia virginiana.These forests occur in the lower elevations (below 2400 feet) of the southern Appalachian Mountains on ridges and upper slopes,typically with southern to western exposures.Environment: These forests occur in the lower elevations (below 2400 feet) of the southern Appalachian Mountains on ridges andupper slopes, typically with southern to western exposures.Vegetation of <strong>Sumter</strong> <strong>National</strong> <strong>Forest</strong> April 30 2004Copyright © 2004 <strong>NatureServe</strong>53
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1 NatureServe is an international o
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types. However, when necessary, mod
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[Association name = floristic nomin
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Generally 5 or fewer occurrences an
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All scientific names for vascular s
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Summary: A short description of the
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Loucks, O. 1996. 100 years after Co
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BIBLIOGRAPHYALNHP [Alabama Natural
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DeYoung, H. R. 1979. The white pine
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Gettman, R. W. 1974. A floristic su
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Lea, C. 2002a. Vegetation classific
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Pittman, Dr. Albert. Personal commu
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Tobe, J. D., J. E. Fairey, III, and